North American Land Mammal Age Research Articles

New occurrences of mammals from McKay Reservoir (Hemphillian, Oregon)

Abstract Encompassing global cooling, the spread of grasslands, and biogeographic interchanges, the Hemphillian North American Land Mammal Age is an important interval for understanding the factors driving ecological and evolutionary change through time. McKay Reservoir near Pendleton, Oregon is a natural laboratory for analyses of these factors. It is remarkable for its small vertebrate fauna including rodents, bats, turtles, and lagomorphs, but also for its larger mammal fossils like camelids, rhinocerotids, canids, and felids. Despite the importance of the site, few revisions to its faunal list have been published since its original description. We expand on this description by identifying taxa not previously known from McKay Reservoir based on specimens collected during fieldwork and through reidentification of previously collected fossils. Newly identified taxa include the borophagine canid Borophagus secundus (Matthew and Cook, 1909), the camelids Megatylopus Matthew and Cook, 1909 and Pleiolama Webb and Meachen, 2004, a dromomerycid, and the equids Cormohipparion Skinner and MacFadden, 1977 and Pseudhipparion Ameghino, 1904. Specimens previously assigned to Neohipparion Gidley, 1903 and Hipparion de Christol, 1832 lack the features necessary to diagnose these genera, which are therefore removed from the site's faunal list. The presence of Borophagus secundus, Cormohipparion, and Pseudhipparion is especially important, because each occurrence represents a major geographic range extension. This refined understanding of the fauna lays the foundation for future studies of taphonomy, taxonomy, functional morphology, and paleoecology—potentially at the population, community, or ecosystem levels—at this paleobiologically significant Miocene locality.

Ibex Hollow Tuff from ca. 12 Ma supereruption, southern Idaho, identified across North America, eastern Pacific Ocean, and Gulf of Mexico

Abstract The Ibex Hollow Tuff, 12.08 ± 0.03 Ma (40Ar/39Ar), is a widespread tephra layer erupted from the Bruneau-Jarbidge volcanic field of southern Idaho. Tephra from this eruption was deposited across much of western and central North America and adjacent ocean areas. We identified the Ibex Hollow Tuff at Trapper Creek, Idaho, near its eruption site, and at 15 distal sites, from the Pacific Ocean to the Gulf of Mexico, by the chemical composition of its glass shards, using electron-microprobe analysis, instrumental neutron activation analysis, and laser-ablation–inductively coupled plasma–mass spectrometry. By these methods, we distinguished the Ibex Hollow Tuff from overlying and underlying tephra layers near its source and at distal sites. Fluvially reworked Ibex Hollow Tuff ash was transported by the ancestral Mississippi River drainage from the interior of the North American continent to the Gulf of Mexico, where it is present within an ~50-m-thick deposit in marine sediments in the subsurface. The minimum fallout area covered by the ash is ~2.7 million km2, with a minimum volume of ~800 km3, and potential dispersal farther to the north and northeast. The areal distribution for the Ibex Hollow Tuff is similar to that of the Lava Creek B (0.63 Ma) supereruption. The Ibex Hollow Tuff represents a unique chronostratigraphic marker allowing a synoptic view of paleoenvironments at a virtual moment in time across a large terrestrial and marine region. The Ibex Hollow Tuff is also an important marker bed for North American Land Mammal Ages, and it coincides with climatic cooling in the middle to late Miocene documented in marine cores.

A new species of Chrysemys (Emydidae: Deirochelyinae) from the latest Miocene-Early Pliocene of Tennessee, USA and its implications for the evolution of painted turtles

Abstract Chrysemys, commonly known as painted turtles, have the largest native biogeographic range of all North American turtles. The presence of a new species, Chrysemys corniculata sp. nov., in the Late Hemphillian-Early Blancan North American Land Mammal Age (latest Miocene-Early Pliocene) of Tennessee provides further data on the evolution of Chrysemys, deirochelyines and emydids. The new fossil species lies basally in Deirochelyinae and suggests that either Chrysemys represents a basal deirochelyine morphology and is one of the oldest genera in the family, or that similar basal morphologies have evolved multiple times throughout deirochelyine evolution. Its occurrence at the same time as Chrysemys picta, during the Hemphillian-Early Blancan, a time of high biodiversity in emydid turtles, suggests either multiple species of Chrysemys during the Late Hemphillian-Early Blancan (at least one in the mid-west and one farther east), or multiple lineages with basal morphologies during this time. Early fossil deirochelyines occur after the greenhouse conditions of the Eocene and the Mid-Miocene Climatic Optimum. Vicariance led to deirochelyines becoming more speciose, including the occurrence of C. corniculata, after the Mid-Miocene Climatic Optimum, potentially suggesting cooler temperatures aided in the evolution of the subfamily and their speciation during the Hemphillian and into the Early Blancan.

Systematics, ecology, and biochronology of blancan horses from Sonora, northwestern Mexico

We describe a set of dental and postcranial horse material recovered from the locality Los Hornitos, northeastern Sonora, northwestern Mexico. The fossil-bearing unit consists of brown silt with intercalation of fine-grained sandstone and bioturbated incipient soils, set in an alluvial environment and considered Pliocene in age. The comparative study of the available sample indicates the presence of two species: the small-sized hipparionine Nannippus peninsulatus and the medium-sized and primitive equine Equus simplicidens. The isotope composition of δ13C and δ18O in dental enamel tooth indicates a C3–C4 mixed diet for these species and suggests the presence of woodland and open areas in Los Hornitos during the Pliocene. The less negative δ13C values observed in E. simplicidens are related to a more C4 diet, preferring to roam open habitats. The coexistence and optimal resource partitioning between these species are explained, considering their differences in dietary behavior, body size, home range, and/or vagility. The recognized horse species are typical of the Blancan North American Land Mammal Age, and their biochronological information from the southwestern United States and central Mexico establishes the probable age of the Los Hornitos at about 3.9–2.6 Ma, corresponding to the Blancan III or late early Blancan – early late Blancan.

An exquisitely preserved skeleton of Eoarctos vorax (nov. gen. et sp.) from Fitterer Ranch, North Dakota (early Oligocene) and systematics and phylogeny of North American early arctoids (Carnivora, Caniformia)

ABSTRACT An exquisitely preserved male skeleton of an early arctoid, Eoarctos vorax new genus and species, provides a unique window into the origin and early divergence of Carnivora. Recovered from the Fitterer Ranch locality in the early Oligocene (Orellan to Whitneyan North American Land Mammal ages) Brule Formation of southwestern North Dakota (∼32 Ma), the new arctoid offers an opportunity to evaluate the fundamental relationships of the caniform (dog-like) carnivorans. Eoarctos vorax possesses a suite of plesiomorphic characters inherited from its miacid ancestors, making it an ideal model for ancestral arctoids. We present a comprehensive treatment of E. vorax, combining traditional description with photographic documentation, microCT, laser scans, and photogrammetry. Showing its plesiomorphic morphology, Eoarctos vorax is scansorial, somewhat like a modern raccoon, retaining the ability to climb trees and lacking cursorial adaptations present in the early canid Hesperocyon. However, E. vorax shows clear signs of durophagous cranio-dental morphology, presumably for an obligatory diet of mollusks, with frequent damage to shell-crushing premolars, plus associated dental infections. We review several other key North American early arctoids and present total-evidence (nuclear DNA and discrete morphological characters) Bayesian and parsimony analyses of Caniformia phylogeny, including extinct stem taxa plus a living member of all modern families. We recognize an endemic North American ursoid clade, Subparictidae, which includes Eoarctos vorax. We demonstrate the importance of North America as an early cradle of evolution for caniform carnivorans, including early precursors of Canidae, Amphicyonidae, Ursidae, and Pinnipedia.

Maybe So, Maybe Not: Canis lepophagus at Hagerman Fossil Beds National Monument, Idaho, USA

A canid dentary is described from the Pliocene Glenns Ferry Formation at Hagerman Fossil Beds National Monument, south-central Idaho, USA. The specimen possesses traits in alliance with and measurements falling within or exceeding those of Canis lepophagus. The dentary, along with a tarsal IV (cuboid) and an exploded canine come from the base of the fossiliferous Sahara complex within the monument. Improved geochronologic control provided by new tephrochronologic mapping by the U.S. Geological Survey-National Park Service Hagerman Paleontology, Environments, and Tephrochronology Project supports an interpolated age of approximately 3.9 Ma, placing it in the early Blancan North American Land Mammal Age. It is conservatively referred to herein as Canis aff. C. lepophagus with the caveat that it is an early and robust example of that species. A smaller canid, initially assigned to Canis lepophagus and then to Canis ferox, is also known from Hagerman. Most specimens of Canis ferox, including the holotype, were recently reassigned to Eucyon ferox, but specimens from the Hagerman and Rexroad faunas were left as Canis sp. and possibly attributed to C. lepophagus. We agree that these smaller canids belong in Canis and not Eucyon but reject placing them within C. lepophagus; we refer to them here as Hagerman-Rexroad Canis. This study confirms the presence of two approximately coyote-sized canids at Hagerman and adds to the growing list of carnivorans now known from these fossil beds.

Dietary niches of creodonts and carnivorans of the late Eocene Cypress Hills Formation.

Modern North American carnivorous mammal assemblages consist of species from a single clade: the Carnivora. Carnivorans once coexisted with members of other meat-eating clades, including the creodonts (Hyaenodontida and Oxyaenida). Creodonts, however, went extinct in North America during the late Eocene and early Oligocene, potentially due to niche overlap and resource competition with contemporary carnivorans. In this study, we employ a community ecology approach to understand whether the dietary niches of coexisting creodonts and carnivorans overlapped during the late Eocene (Chadronian North American Land Mammal Age), a time when creodonts were dwindling and carnivorans were diversifying. We quantify niche overlap based on inferences of diet from carnassial tooth shape estimated using Orientation Patch Count, Dirichlet’s Normal Surface Energy, and linear dental measurements as well as from body mass for all species in the Calf Creek Local Fauna of Cypress Hills, Saskatchewan (Treaty 4 land). Although creodonts and carnivorans shared characteristics of their carnassial tooth shape, suggesting similar chewing mechanics and feeding habits, we find that marked differences in body size likely facilitated niche partitioning, at least between the largest creodonts and carnivorans. Calculations of prey focus masses and prey mass spectra indicate that only the smallest creodont may have experienced significant competition for prey with the coeval carnivorans. We suggest that the ultimate extinction of creodonts from North America during the late Eocene and Oligocene was unlikely to have been driven by factors related to niche overlap with carnivorans.

Small Colubroids from the Late Hemphillian Gray Fossil Site of Northeastern Tennessee

The Gray Fossil Site is a rich Hemphillian (North American Land Mammal Age) locality in northeastern Tennessee, USA, and has produced tens of thousands of fossils of multiple taxa including hundreds of individual snake skeletal remains. One of only five sites of similar age east of the Mississippi River, the Gray Fossil Site is crucial to our understanding of snake evolution and diversity. Microfauna were collected by screenwashing sediment and picking under a microscope. Specimens were identified using a suite of measurements and characters. Analyzed here are vertebral fossils identified as belonging to various colubrid taxa including Worm Snakes (Carphophis sp.), Ring-Necked Snakes (Diadophis sp.), Hook-Nosed Snakes (Gyalopion sp.), and Hognose Snakes (Heterodon meadi sp. nov.). Carphophis sp. and Gyalopion sp. represent first occurrences in the Mio-Pliocene and dramatically extend their temporal range from the Pleistocene. The four snakes, including one new species, are in accordance with the previously identified taxa by Jasinski and Moscato (2017; Journal of Herpetology 5:245–257), represent a modern assemblage of advanced snakes by the Mio-Pliocene and support a forested environment in northeastern Tennessee.

Geometric morphometrics and anatomical network analyses reveal ecospace partitioning among geoemydid turtles from the Uinta Formation, Utah.

We present new fossil records of the geoemydid turtle Bridgeremys pusilla from the Uinta Formation of Utah. Turtles are abundant throughout the unit, and known taxa are similar to those from the older strata in the Upper Green River Basin in Wyoming from the Bridger and Washakie Formations. B. pusilla is known from Bridgerian deposits but was not previously known from after the Turtle Bluff Member of the Bridger Formation. The taxon was coveal with two species of the geoemydid Echmatemys (E. callopyge and E. wyomingensis), a common genus of extinct pond turtles known primarily from lacustrine and fluvial deposits in western North America, including the Uinta Basin. In addition to previously documented morphological differences, our geometric morphometric analyses revealed significant differences in epiplastral morphology between B. pusilla and the two coeval Echmatemys species. Bridgeremys pusilla shared several morphological characters with Testudinidae. However, our anatomical network analysis suggests that the carapace of B. pusilla distributed stress forces in a manner more similar to emydids (basal and derived) than to derived testudinoids (Testudinidae and Emydidae), including Echmatemys species. This finding changes our understanding of the ecology of the species and sheds light onto how geoemydid turtles of the Uinta Formation may have partitioned the available ecospace. These new Uintan records extend the geographic range of B. pusilla into the Uinta Basin and stratigraphically through the top of the Uinta Formation, extending the temporal range of the taxon by more than 4 million years through the Uintan North American Land Mammal Age to the base of the Duchesne River Formation.

Magnetostratigraphy of lower Paleocene strata in the Ferris Formation, Hanna Basin, Wyoming, with refined resolution of the Pu1–Pu2 interval-zone boundary of the Puercan North American Land Mammal Age

ABSTRACT Recent studies of early Paleocene stratigraphic sections across the U.S. Western Interior are refining our understanding of the biotic recovery in the aftermath of the Cretaceous–Paleogene (K–Pg) mass extinction event. Herein we present magnetostratigraphic data from an approximately 600-m-thick section of strata of the Ferris Formation in the Hanna Basin, south-central Wyoming, that spans the K–Pg boundary and includes in conformable superposition the three subdivisions of the earliest Paleocene Puercan North American Land Mammal Age: interval zones Pu1, Pu2, and Pu3. Prior studies on early Paleocene stratigraphic sections in Montana to the north of Hanna Basin and in Colorado and New Mexico to the south have been only marginally successful in defining the temporal boundaries of these divisions. This earlier work typically restricted Pu1 entirely within reversed magnetochron C29r, Pu2 entirely within normal magnetochron C29n, and at least the early part of Pu3 in C29n, as well. Results of the present study confirm what has been only tentatively suggested previously: that interval zone Pu2 begins in the youngest part of C29r, with later Pu2 fauna extending into C29n. Although Pu3 is known to begin in C29n, its younger limit remains unknown in our Hanna Basin section, because the top of the local Puercan is well above the stratigraphic level of our sampling for this project. We estimate a date for the Pu1–Pu2 boundary in the Hanna Basin section of approximately 65.82 Ma—nearly 70 k.y. earlier than prior estimates. This boundary marks the transition from the Pu1 survival fauna to later phases of the mammalian recovery characterized by a significant increase in diversity, appearance of considerably larger-bodied mammals, and greater dental and dietary specializations.

A latest Eocene (Chadronian) brontothere (Mammalia, Perissodactyla) from the Antero Formation, South Park, Colorado

ABSTRACTLate Eocene brontotheres are documented most prevalently from formations in the Great Plains of North America. Here we describe UCM 109045, a mandible and lower dentition of a brontothere recovered from a latest Eocene (Chadronian) locality in the Antero Formation in South Park, Colorado. This is a high-altitude locality in which vertebrate fossils are rare. Lower incisor number and presence of a long postcanine diastema indicate that UCM 109045 does not belong to Megacerops coloradensisLeidy, 1870, by far the most abundant brontothere from the Chadronian North American Land Mammal Age. Instead, UCM 109045 is morphologically most similar to Protitanops curryiStock, 1936, from the early Chadronian of the southwestern United States, and nomen dubium Megacerops primitivusLambe, 1908, from the Chadronian of Saskatchewan, Canada. It is possible that Megacerops kuwagatarhinusMader and Alexander, 1995, is a junior synonym of M. primitivus. If UCM 109045 belongs to Megacerops primitivus (= M. kuwgatarhinus), it would support the hypothesis that only two species of brontothere—M. primitivus (= M. kuwgatarhinus) and M. coloradensis—survived into the latest Eocene. Regardless of its exact identification, the discovery of UCM 109045 in the Antero Formation provides insight into a poorly understood, high-altitude locality in North America from just before brontothere extinction at the Eocene–Oligocene boundary.

New specimens of the mesonychid Dissacus praenuntius from the early Eocene of Wyoming and evaluation of body size through the PETM in North America

The Mesonychia is a group of archaic carnivorous mammals of uncertain phylogenetic affinities with a Holarctic distribution during the Paleogene. Intensive fossil collecting efforts in the Bighorn Basin, Wyoming, have resulted in recovery of the largest sample and most complete specimens yet known of the mesonychid Dissacus praenuntius from the second biozone of the Wasatchian North American Land Mammal Age (Wa-0). The Wa-0 biozone corresponds to the body of the Paleocene-Eocene Thermal Maximum (PETM), a brief but intense global warming event that occurred ~56myr ago that significantly impacted terrestrial mammal faunas, including dwarfing in many mammal lineages. To evaluate the potential response of this lineage to climate change, we compared the PETM sample of D. praenuntius with those recovered from just before the PETM in the last biozone of the Clarkforkian North American Land Mammal Age (Cf-3) and just after the PETM in the Wa-1 biozone. While the sample size is still too small to say with certainty, tooth size (as a proxy for body weight) of D. praenuntius appears to be smaller during the late PETM than during either the pre-PETM Cf-3, or post-PETM Wa-1 biozones, suggesting the possibility of a muted dwarfing response to the PETM. However, the pattern observed for D. praenuntius differs from that of many other PETM mammals, as the shift to smaller body size is less pronounced and may have only occurred in late Wa-0.

Paleobiology of a large mammal community from the late Pleistocene of Sonora, Mexico

AbstractA paleontological deposit near San Clemente de Térapa represents one of the very few Rancholabrean North American Land Mammal Age sites within Sonora, Mexico. During that time, grasslands were common, and the climate included cooler and drier summers and wetter winters than currently experienced in northern Mexico. Here, we demonstrate restructuring in the mammalian community associated with environmental change over the past 40,000 years at Térapa. The fossil community has a similar number of carnivores and herbivores whereas the modern community consists mostly of carnivores. There was also a 97% decrease in mean body size (from 289 kg to 9 kg) because of the loss of megafauna. We further provide an updated review of ungulates and carnivores, recognizing two distinct morphotypes ofEquus, includingE.scottiand a slighter species; as well asPlatygonus compressus;Camelops hesternus;Canis dirus; andLynx rufus;and the first regional records ofPalaeolama mirifica,Procyon lotor, andSmilodoncf.S.fatalis. The Térapa mammals presented here provide a more comprehensive understanding of the faunal community restructuring that occurred in northern Mexico from the late Pleistocene to present day, indicating further potential biodiversity loss with continued warming and drying of the region.

Rodents (Mammalia) from the Troublesome Formation, Late Oligocene to Miocene (Middle Arikareean–Early Clarendonian) of Colorado

The Troublesome Formation of northcentral Colorado ranges in age from the middle Arikareean North American Land Mammal Age (NALMA: latest Oligocene) to the early Clarendonian NALMA (early–late Miocene), a range of 14.5 million years. The rodent fauna from this formation varies greatly through the section, which is dominated by entoptychine geomyids in the Arikareean, mylagaulids in the Hemingfordian, and heteormyids in the Barstovian. The Clarendonian is the least-well represented horizon from which only four species are recognized. Of the nearly forty species of rodents identified, there is only one new genus, Argaleogaulus a primitive promylagauline mylagaulid from the Arikareean. There are 12 new species recognized: the mylagaulid Argaleogaulus primoticus; a sciurid, Protospermophilus parvus; six heteromyids, Harrymys cyanothos, Harrymys taussigi, Schizodontomys bareia, Balantiomys coloradensis, Oregonomys perilaccos, and Cupidinimus robinsoni; and four entoptychine geomyids, Pleurolicus compressus, Pleurolicus mensae, Gregorymys montanus, and Entoptychus rensbergeri. The genera from the Troublesome Formation are similar to those of equivalent faunas throughout North America. The presence of the unique species of otherwise common North American genera suggests some minor degree of isolation of the basin during the time of formation.

Taxonomic Revision of the Ischyromyid Rodent Thisbemys from the Uintan (Middle Eocene) of Utah Based on New Previously Undescribed Material

The ischyromyid rodent Thisbemys Wood, 1959, is known from the lower Eocene (Wasatchian North American Land Mammal Age [ = NALMA]) through the middle Eocene (Uintan NALMA) of western North America....

Additional records and stratigraphic distribution of the middle Eocene carettochelyid turtle Anosteira pulchra from the Uinta Formation of Utah, North America.

BackgroundAnosteira pulchra is one of two species of the obligately-aquatic freshwater clade Carettochelyidae (pig-nosed turtles) from the Eocene of North America. Anosteira pulchra is typically rare in collections, and their distribution is poorly documented. The Uinta Formation [Fm.] contains a diverse assemblage of turtles from the Uintan North American Land Mammal Age. Whereas turtles are abundantly preserved in the Uinta Fm., A. pulchra has been reported only from a few specimens in the Uinta C Member.MethodsWe describe new records of Anosteira pulchra from the Uinta Basin and analyze the distribution of 95 specimens from multiple repositories in the previously published stratigraphic framework of the middle and upper Uinta Fm.ResultsHere we report the first records of the species from the Uinta B interval, document it from multiple levels within the stratigraphic section and examine its uncommon appearance in only approximately 5% of localities where turtles have been systematically collected. This study details and extends the range of A. pulchra in the Uinta Fm. and demonstrates the presence of the taxon in significantly lower stratigraphic layers. These newly described fossils include previously unknown elements and associated trace fossils, with new anatomical information presented. This study provides insight into the taxonomy of Anosteira spp. in the middle Eocene, and suggests the presence of a single species, though no synonymy is defined here due to limits in Bridger material.

Marsupials (Mammalia, Metatheria) from the Brule Formation (Whitneyan, Oligocene) North Dakota

AbstractFive marsupial species are recognized from the Brule Formation at two localities in southwestern North Dakota: Fitterer Ranch and Obritsch Ranch (middle Oligocene; Whitneyan North American Land Mammal Age [NALMA]). The herpetotheriidsHerpetotherium fugaxCope, 1873a,Copedelphys superstesnew species, and the peradectidNanodelphys hunti(Cope, 1873b) are represented at both localities. A fourth species isH. sp., cf.H.merriami(Stock and Furlong, 1922), represented by a single specimen from Fitterer Ranch, being limited elsewhere to the later Arikareean NALMA. A fifth species is represented by two isolated lower cheek teeth, interpreted as m1s, from Fitterer Ranch that are unique in lacking a trigonid (only two cusps present) while having a well-developed talonid. These specimens are referred to an indeterminate herpetotheriine species. The new species ofCopedelphysis distinct from other species of the genus in that the anterior two lower molars are enlarged relative to the posterior molars. Overall, this new species is more similar in proportions to the latest Eocene (Chadronian)C.titanelix(Matthew, 1903) than the Oligocene (Orellan and Whitneyan)C.stevensoni(Cope, 1873b). This study adds a third and fourth Whitneyan marsupial fauna from the Great Plains region of North America, increases the known diversity of Whitneyan marsupials, and provides further evidence that marsupial diversity during the late Paleogene in North America was relatively stable until the late early Arikareean NALMA.UUID:http://zoobank.org/b8534802-b9ed-4120-baca-fc72917f7d6a

REIDENTIFICATION OF LATE MIDDLE EOCENE “UINTACYON” FROM THE GALISTEO FORMATION (NEW MEXICO, U.S.A.) AS AN EARLY BEARDOG (MAMMALIA, CARNIVORA, AMPHICYONIDAE)

Museum of Comparative Zoology (MCZ) VPM-4225, a carnivoran dentary, is described here for the first time, 82 years after it was collected in the Duchesnean (late middle Eocene) portion of the Galisteo Formation in New Mexico. It evidently corresponds to the specimen—never identified by its catalog number, never described, and considered missing for the last 41 years—that formed the basis for the purported occurrence of the early carnivoraform Uintacyon in the formation reported in 1943. The specimen is here reidentified as one of the earliest amphicyonids, Angelarctocyon, which was previously known from only two specimens from the Chambers Tuff of Texas. With this reidentification, there is no longer a valid occurrence of Uintacyon postdating the Uintan North American Land Mammal Age, adding to the evidence for rapid replacement of ancient carnivoramorphan lineages by early crown-clade carnivorans in the late middle Eocene, across a period of severe climatic fluctuations 42–40 million years ago.

New material of fossil rodents (Mammalia) from the Eocene (Bridgerian-Uintan) Washakie Formation, southcentral Wyoming

Eight species of rodents are described based on previously unreported fossil material from the early Eocene Adobe Town Member of the Washakie Formation of Wyoming. Among these are two new species: the sciuravid Pauromys turnbulli and the cylindrodont Pareumys flynni. The overall rodent fauna consists of twelve species. The greatest change in the rodent fauna is between the lower and middle units of the Adobe Town Member; six species having their last occurrence in the lower unit and three species having their first occurrence in the middle unit. This coincides with the boundary of the Bridgerian and Uintan North American Land Mammal Ages.

Fallout tuffs in the Eocene Duchesne River Formation, northeastern Utah—ages, compositions, and likely source

Thin fallout tuffs are common in the terrestrial deposits of the Eocene Duchesne River Formation on the flanks of the Uinta Mountains of eastern Utah. Their ages and compositions provide new insight into the tectonic events and magmatic history of the western Cordillera and provide important constraints on the Cenozoic land mammal chronology. Whole-rock compositions of the volcanic ash show that they underwent post-emplacement argillic alteration, typical of a wetland/floodplain depositional setting. However, immobile element ratios and abundances, such as Zr/Ti, La/Nb, and Y are typical of rhyolites formed in a subduction-related setting. Glass shards preserved in one sample all had SiO2 values >75%, typical of high-silica rhyolite. Preserved phenocrysts in the ash beds include quartz, sanidine, plagioclase, and biotite with variable amounts of accessory zircon, apatite, titanite, and allanite. Biotite compositions have Fe/(Fe+Mg) ratios typical of calc-alkaline igneous rocks and clusters of chemical compositions suggest a genetic relationship to three or four separate eruptions. Sanidine compositions from five samples range from Or73 and Or79. Only one sample had preserved plagioclase with compositions ranging between An22 – An49. Allanite from the ash beds has lower total rare earth elements (REE) concentrations than allanite from other well-studied rhyolites. Titanite in one sample has lower concentrations of REE, Fe, and Al than expected of rhyolites and is probably detrital.Plagioclase and sanidine from two different tuff beds near the middle of the Duchesne River Formation yielded analytically indistinguishable 40Ar/39Ar ages of 39.47 ± 0.16 Ma and 39.36 ± 0.15 Ma, respectively. These dates, along with the compositional data seem to limit the eruptive source for these fallout tuffs to the northeast Nevada volcanic field, one of the few volcanically active regions of western North America at the time. These new radiometric ages, along with stratigraphic relations and previously published ages for tuffs in the Bishop Conglomerate (which unconformably overlies the Duchesne River Formation), constrain the timing of late Laramide uplift in the region from 39 to about 37 Ma and post-Laramide epeirogenic uplift from 34 Ma to 30 Ma. Finally, the ages also provide additional evidence that the Duchesnean North American Land Mammal Age ended in the Eocene, which was originally named and defined from the Duchesne River Formation.