North American Holstein-Friesian Research Articles

Intensification strategies for temperate hot-summer grazing dairy systems in South America: Effects of feeding strategy and cow genotype

Pasture-based dairy systems present the opportunity to increase productivity per hectare through increasing stocking rate and forage utilization. However, in the temperate hot-summer region of South America, different productive strategies are being adopted by farmers. The aim of this study was to quantify the effect of feeding strategy (FS) and cow genotype (G) on individual animal and whole-farm biophysical performance. A design with 2 × 2 levels of intensification aiming to increase home-grown forage utilization and milk output per hectare was evaluated. The experiment was a randomized complete block design with a 2 × 2 factorial arrangement of treatments, combining 2 feeding strategies with varying proportions of grazing in the annual feeding budget [grass fixed (GFix) and grass maximum (GMax)] and 2 Holstein Friesian cow genotypes [New Zealand (NZHF) or North American Holstein Friesian (NAHF)]. The effects of FS, G, and their interaction were analyzed using mixed models. New Zealand Holstein Friesian cows presented lower individual milk yield and higher milk component concentrations, maintained higher average body condition score, and increased body weight (BW) throughout the experiment, while presenting a better reproductive performance compared with the NAHF cows. Although all farmlets were planned at the same stocking rate on a per kilogram of BW basis, the current stocking rate changed as a result of animal performance and grass utilization resulting in NZHF cows achieving greater BW per hectare. The superior stocking rate led to greater milk solids production and feed consumption per hectare for the systems with NZHF cows. The GFix feeding strategy resulted in greater total home-grown forage harvest and conserved forage surplus than GMax. Overall, it was feasible to increase stocking rate and increase milk production per hectare from home-grown forage with differing feeding strategies and Holstein Friesian cow genotypes within grazing systems located in the temperate hot-summer climate region of South America. The interactions reported between FS × G highlight the superior productivity per hectare of NZHF cows within the GMax feeding strategy based on maximizing grazed pasture, which could represent a competitive intensification strategy in terms of cost of production for this region.

Whole-farm modelling of grazing dairy systems in Uruguay

CONTEXTModelling grazing dairy systems from the temperate hot-summer climate region of South America is challenging due to the absence of suitable whole-farm models developed or evaluated in this region. The use of whole-farm models developed in other regions represents an opportunity. However, the accuracy and precision of their predictions need to be assessed. OBJECTIVESThe present study evaluated the predictive ability of Farmax Dairy Pro, a whole-farm model developed in New Zealand, to simulate grazing dairy systems in Uruguay. METHODSData used for the model evaluation was obtained from a dairy farmlet study carried out in Uruguay. The study aimed to explore four intensification strategies based on increasing home-grown forage utilisation and milk output per hectare. The field experiment consisted of four farmlets using two feeding strategies: [Grass Maximum (GMAX) and Grass Fixed (GFIX), based on the amount of grazed herbage in the diet], and two cow genotypes [New Zealand (NZHF) or North American Holstein-Friesian (NAHF)]. The four farmlets were modelled with Farmax for two consecutive years. Model evaluation was performed using standard regression, dimensionless and error index statistics. The model was evaluated by comparing predicted versus observed monthly patterns of milk, milk fat, milk protein and milk solids (MS; milk fat + milk protein) yields, body condition score (BCS), body weight (BW), DM intake (DMI) and net pasture growth rate (PGR). An application of the model was demonstrated by modelling three scenarios for the GMAX-NZHF farmlet. RESULTS AND CONCLUSIONSThe predictive ability of Farmax was similar for the four farmlets modelled, including patterns over time, for all the variables evaluated. The model provided a robust prediction for monthly patterns of milk and milk components yields at a herd level for total DMI and PGR. The model had a moderate ability to predict monthly patterns of individual milk and milk components yields and BCS, and a poor ability to predict BW. The scenario modelling results indicate that the model could be used with confidence to simulate different farm system alternatives. Overall, the Farmax Dairy Pro model had the potential to provide adequate predictions for grazing dairy systems from Uruguay. SIGNIFICANCEThis model will allow the exploration of future intensification pathways for grazing dairy systems in Uruguay and the region, including changes in the forage sequence, stocking rate and calving season. Further adjustments of the model will expand the range of systems and latitudes for this model to be utilised.

The effect of stocking rate and calving date on milk production of Holstein–Friesian dairy cows

The objective of this study was to evaluate the effect of stocking rate (SR) and calving date (CD) on milk production, BW and BCS within grass-based production systems post-European Union milk quotas using modern grazing management practices and high genetic potential Holstein–Friesian (HF) spring calving dairy cattle over a two year period. Two groups of HF dairy cows with different mean CD were established from within the existing research herd at Moorepark (Teagasc, Ireland). Animals were assigned either to an early calving (mean CD: 12 February) treatment or a late calving (mean CD: 25 February) treatment. Animals within each CD treatment were randomly allocated to 1 of 3 SR treatments, Low (2.51 cows/hectare (ha), Medium (2.92cows/ha) and High (3.28cows/ha) which were designed to represent 3 alternative whole farm SR in a post-European Union milk quota, spring calving, grass-based milk production system. A total of 138 spring-calving dairy cows, comprised of two strains of HF (North American HF and New Zealand HF genetic strains), were used during 2009 and 2010, respectively. The effects of CD, SR treatment, genetic strain and their interactions on milk production per cow and per ha, body weight and body condition score were analyzed. Although reducing per animal production, increased SR resulted in increased milk and milk solids production per ha. The results also indicate that although CD had no effect on total lactation performance, adjusting mean CD may be an effective strategy to align animal requirements and grass supply and reduce the requirement for supplements at increased SR in early lactation.

Calculation of a cow culling merit index including specific heterosis in a multibreed dairy population

Abstract. The objective of this study was to compare two models for the estimation of producing values (EPV) for lactation yields of milk, fat and protein, and calving interval (CI), which were combined in an index called the Cow Culling Merit Index (CMI), in Irish dairy cattle. Data comprised 188 927 records for production and 157 117 records for CI, collected on North American Holstein Friesian (HO), Friesian (FR), Jersey (JE), and Montbéliarde (MO) pure breeds, and some of their crosses. Cows calved from 2002 to 2006 and were from parities 1 to 5. Coefficients of specific heterosis for HO×FR, HO×JE, and HO×MO were calculated for each cow from parental breed information. The coefficient of general heterosis (GH) for each cow was obtained as the sum of the specific coefficients previously estimated. Model 1 included fixed effects of contemporary group, age at calving within parity, linear regression on gene proportions for FR, JE, and MO, and linear regression on the coefficient of expected GH. Additive genetic, permanent environmental, and error were random effects. Model 2 was based on Model 1 but GH was replaced by linear regressions on coefficients of expected specific heterosis for HO×FR, HO×JE, and HO×MO. Estimated producing values were calculated as the sum of estimated breeding value, permanent environmental and heterosis effects. The inclusion of coefficients of specific heterosis in the model did not produce re-ranking of animals but important differences in EPVs were observed in crossbred cows. These changes are important if EPVs are used to develop a culling merit index.

Dry matter intake and feed efficiency profiles of 3 genotypes of Holstein-Friesian within pasture-based systems of milk production

The primary objective of the study was to quantify the effect of genetic improvement using the Irish total merit index (Economic Breeding Index) on dry matter intake and feed efficiency across lactation and to quantify the variation in performance among alternative definitions of feed efficiency. Three genotypes of Holstein-Friesian dairy cattle were established from within the Moorepark dairy research herd: 1) low Economic Breeding Index North American Holstein-Friesian representative of the Irish national average dairy cow, 2) high genetic merit North American Holstein-Friesian, and 3) high genetic merit New Zealand Holstein-Friesian. Animals from within each genotype were randomly allocated to 1 of 2 possible intensive pasture-based feed systems: 1) the Moorepark pasture system (2.64 cows/ha and 500kg of concentrate supplement per cow per lactation) and 2) a high output per hectare pasture system (2.85 cows/ha and 1,200kg of concentrate supplement per cow per lactation). A total of 128 and 140 spring-calving dairy cows were used during the years 2007 and 2008, respectively. Each group had an individual farmlet of 17 paddocks, and all groups were managed similarly throughout the study. The effects of genotype, feed system, and the interaction between genotype and feed system on dry matter intake, milk production, body weight, body condition score, and different definitions of feed efficiency were studied using mixed models with factorial arrangements of genotypes and feed systems accounting for the repeated cow records across years. No significant genotype-by-feed-system interactions were observed for any of the variables measured. Results showed that aggressive selection using the Irish Economic Breeding Index had no effect on dry matter intake across lactation when managed on intensive pasture-based systems of milk production, although the ranking of genotypes for feed efficiency differed depending on the definition of feed efficiency used. Performance of animals grouped on alternative definitions of feed efficiency showed that conventional definitions such as feed conversion efficiency or residual feed intake may be inappropriate measures of efficiency for lactating dairy cows. An alternative definition, residual solids production, is proposed. This definition of feed efficiency identifies animals that produce greater volumes of milk solids at similar levels of feed intake without excessive body tissue mobilization and with improved fertility performance. The results also suggest that although there are differences in feed efficiency between strains of Holstein-Friesian, there is also variation within genotypes so that improvements in feed efficiency can be realized if the appropriate definition of feed efficiency is incorporated into breeding programs.

Crossbreeding effects on milk yield traits and calving interval in spring‐calving dairy cows

The aim of the study was to assess crossbreeding effects for 305-day milk, fat, and protein yield and calving interval (CI) in Irish dairy cows (parities 1 to 5) calving in the spring from 2002 to 2006. Data included 188,927 records for production traits and 157,117 records for CI. The proportion of genes from North American Holstein Friesian (HO), Friesian (FR), Jersey (JE) and Montbéliarde (MO) breeds, and coefficients of expected heterosis for HOxFR, HOxJE and HOxMO crosses were calculated from the breed composition of cows' parents. The model used to assess crossbreeding effects accounted for contemporary group, age at calving within parity, linear regression on gene proportions for FR, JE and MO, and linear regression on coefficients of expected heterosis for HOxFR, HOxJE and HOxMO, as fixed effects, and additive genetic, permanent environmental and residual as random. Breed effects for production traits were in favour of HO, while for CI were in favour of breeds other than HO. The highest heterosis estimates for production were for HOxJE, with first-generation crosses yielding 477 kg more milk, 25.3 kg more fat, and 17.4 kg more protein than the average of the parental breeds. The highest estimate for CI was for HOxMO, with first-generation crosses showing 10.2 days less CI than the average of the parental breeds. Results from this study indicate breed differences and specific heterosis effects for milk yield traits and fertility exist in Irish dairy population.

The influence of genetic selection and feed system on the reproductive performance of spring-calving dairy cows within future pasture-based production systems

Three genetic groups of Holstein-Friesian dairy cows were established from within the Moorepark (Teagasc, Ireland) dairy research herd: LowNA, indicative of the Irish national average-genetic-merit North American Holstein-Friesian; HighNA, high-genetic-merit North American Holstein-Friesian; HighNZ, high-genetic-merit New Zealand Holstein-Friesian. Genetic merit in this study was based on the Irish total merit index, the Economic Breeding Index. Animals from within each genetic group were randomly allocated to 1 of 2 possible post-European Union-milk-quota pasture-based feeding systems (FS): 1) The Moorepark (MP) pasture system (2.64 cows/ha and 500kg of concentrate supplement per cow per lactation) and 2) a high output per hectare (HC) pasture system (2.85 cows/ha and 1,200kg of concentrate supplement per cow per lactation). A total of 126, 128, and 140 spring-calving dairy cows were used during the years 2006, 2007, and 2008, respectively. Each group had an individual farmlet of 17 paddocks, and all groups were managed similarly throughout the study. The effects of genetic group, FS, and the interaction between genetic group and FS on reproductive performance, body weight, body condition score, and blood metabolite concentrations were studied using mixed models with factorial arrangements of genetic groups and FS. Odds ratios were used in the analysis of binary fertility traits, and survival analysis was used in the analysis of survival after first calving. When treatment means were compared, the HighNA and HighNZ genotypes (with greater genetic merit for fertility performance) had greater first-service pregnancy rates and had a greater proportion of cows pregnant after 42 d of the breeding season than the LowNA group. Both HighNA and HighNZ genotypes were submitted for artificial insemination earlier in the breeding season and had greater survival than the LowNA genotype. There was no significant FS or genotype by FS interactions for any of the reproductive, blood metabolite, body weight, or body condition score measures. The results demonstrate that increased genetic merit for fertility traits resulted in improved reproductive performance and that the poor reproductive capacity of inferior-genetic-merit animals for fertility was not improved through concentrate supplementation at pasture.

Differences in the expression of genes involved in the somatotropic axis in divergent strains of Holstein-Friesian dairy cows during early and mid lactation

Differences in genetic selection criteria for dairy cows internationally have led to divergence in the Holstein-Friesian breed. The objective of this study was to compare hepatic expression of genes of the somatotropic axis in the North American Holstein-Friesian and the New Zealand Holstein-Friesian strains of dairy cow at early and mid lactation. Mature cows of both the North American Holstein-Friesian (n=10) and New Zealand Holstein-Friesian (n=10) strains were selected. Liver tissue was collected by percutaneous punch biopsy from all cows at 35 and 140 d postpartum, representing early and mid lactation, respectively. Total RNA was extracted and the hepatic expression of genes involved in the control of the somatotropic axis was examined. Abundance of insulin-like growth factor (IGF)-1 mRNA was greater in the New Zealand strain, concomitant with a tendency for increased expression of acid-labile subunit mRNA. Across strains, mRNA abundance of IGF-binding protein-1, IGF-binding protein-2, and growth hormone receptor 1A decreased from d 35 to 140 postpartum, whereas expression of IGF-1 and acid-labile subunit tended to increase. Abundance of suppressor of cytokine signaling-3 mRNA was increased at d 140 postpartum. Both the strain of Holstein-Friesian cow and the stage of lactation influenced expression of genes controlling the somatotropic axis in hepatic tissue.

Extending lactation in pasture-based dairy cows. II: Effect of genetic strain and diet on plasma hormone and metabolite concentrations

Fifty-six genetically divergent New Zealand and North American Holstein-Friesian (HF) cows grazed pasture, and were offered 0, 3, or 6kg of concentrate DM/cow per day for an extended lactation (605±8.3 d in milk; mean±standard error of the mean). Weekly blood samples collected from individual cows from wk 1 to 10 postpartum (early lactation), and from wk 47 to 63 postpartum (extended lactation) were analyzed for nonesterified fatty acids (NEFA), glucose, insulin, leptin, growth hormone (GH), insulin-like growth factor-I (IGF-I), calcium, and urea. During early lactation, NEFA and GH concentrations were greater and IGF-I concentrations were less, and increased at a slower rate in North American HF. During this 10-wk period, there were no strain effects on plasma glucose, leptin, insulin, or calcium. During the extended lactation period, North American HF had greater NEFA and GH concentrations; there were strain×diet interactions for insulin and leptin, and a tendency for a strain×diet interaction for glucose. These interactions were primarily due to greater plasma insulin, leptin, and glucose concentrations in the New Zealand HF fed 6kg of concentrate DM/cow per day, a result of excessive body condition in this treatment. In this period, there was no strain effect on plasma IGF-I, calcium, or urea concentration. During early lactation, there was a linear increase in glucose and IGF-I, and a linear decrease in GH and urea with increasing concentrate in the diet. However, plasma calcium, NEFA, insulin, and leptin remained unchanged. During the extended lactation period, there was an effect of feed supplementation on GH and urea, which decreased linearly with increasing concentrate in the diet. There was, however, no supplementation effect on NEFA, calcium, or IGF-I. These data indicate potential strain differences in recoupling of the somatotropic axis, insulin resistance, and energy partitioning, and may help explain the physiology behind the previously reported greater milk production and body condition score loss in North American HF. The results have implications for breeding and diet management during an extended lactation.

A comparison of energy balance and metabolic profiles of the New Zealand and North American strains of Holstein Friesian dairy cow

The milk production, energy balance (EB), endocrine and metabolite profiles of 10 New Zealand Holstein Friesian (NZ) cows and 10 North American Holstein Friesian (NA) cows were compared. The NA cows had greater peak milk yields and total lactation milk yields (7387 v. 6208 kg; s.e.d. = 359), lower milk fat and similar protein concentrations compared with the NZ cows. Body weight (BW) was greater for NA cows compared with NZ cows throughout lactation (596 v. 544 kg; s.e.d. = 15.5), while body condition score (BCS) tended to be lower. The NA strain tended to have greater dry matter intake (DMI) (17.2 v. 15.7 kg/day; s.e.d. = 0.78) for week 1 to 20 of lactation, though DMI as a proportion of metabolic BW was similar for both strains. No differences were observed between the strains in the timing and magnitude of the EB nadir, interval to neutral EB, or mean daily EB for week 1 to 20 of lactation. Plasma concentrations of glucose and insulin were greater for NA cows during the transition period (day 14 prepartum to day 28 postpartum). Plasma IGF-I concentrations were similar for the strains at this time, but NZ cows had greater plasma IGF-I concentration from day 29 to day 100 of lactation, despite similar calculated EB. In conclusion, the results of this study do not support the premise that the NZ strain has a more favourable metabolic status during the transition period. The results, however, indicate that NZ cows begin to partition nutrients towards body reserves during mid-lactation, whereas NA cows continue to partition nutrients to milk production.

Extending Lactation in Pasture-Based Dairy Cows: I. Genotype and Diet Effect on Milk and Reproduction

The aim of this study was to test the feasibility of extended lactations in pastoral systems by using divergent dairy cow genotypes [New Zealand (NZ) or North American (NA) Holstein-Friesian (HF)] and levels of nutrition (0, 3, or 6 kg/d of concentrate dry matter). Mean calving date was July 28, 2003, and all cows were dried off by May 6, 2005. Of the 56 cows studied, 52 (93%) were milking at 500 d in milk (DIM) and 10 (18%) were milking at 650 DIM. Dietary treatments did not affect DIM (605±8.3; mean±SEM). Genotype by diet interactions were found for total yield of milk, protein, and milk solids (fat + protein), expressed per cow and as a percentage of body weight. Differences between genotypes were greatest at the highest level of supplementation. Compared with NZ HF, NA HF produced 35% more milk, 24% more milk fat, 25% more milk protein, and at drying off had 1.9 units less body condition score (1 to 10 scale). Annualized milk solids production, defined as production achieved during the 24-mo calving interval divided by 2 yr, was 79% of that produced in a normal 12-mo calving interval by NZ HF, compared with 94% for NA HF. Compared with NZ HF, NA HF had a similar 21-d submission rate (85%) to artificial insemination, a lower 42-d pregnancy rate (56 vs. 79%), and a higher final nonpregnancy rate (30 vs. 3%) when mated at 451 d after calving. These results show that productive lactations of up to 650 d are possible on a range of pasture-based diets, with the highest milk yields produced by NA HF supplemented with concentrates. Based on the genetics represented, milking cows for 2 yr consecutively, with calving and mating occurring every second year, may exploit the superior lactation persistency of high-yielding cows while improving reproductive performance.

Trends in milk production, calving rate and survival of cows in 14 Irish dairy herds as a result of the introgression of Holstein-Friesian genes

AbstractTrends in milk production, calving rates, and survival were monitored on a potential 5580 primiparous and multiparous Holstein-Friesian dairy cows across 14 Irish seasonal spring-calving dairy farms between the years 1990 and 2001. Over this period calving rate to first service (CALV1) reduced by 0·96% per year (55 to 44%;P< 0·001), calving rate to first and second service (CALV12) reduced by 0·84% per year ( 77 to 70%;P< 0·001) and herd average parity number reduced by 0·10 lactation per year (4·3 to 3·5;P<0·001). The proportion of North American Holstein Friesian (NAHF) genes in the cows increased by 5·5% per year (8 to 63%;P<0·001), while pedigree index for milk yield (PIMILK) of the cows increased by 25 kg per year (P<0·001). The predicted difference of the sires of the cows for calving interval and survival increased by 0·5 days (P<0·001) and reduced by 0·12% (P<0·001) per year, respectively. A negative association was found between increased phenotypic milk yield, NAHF and PIMILKand reduced calving rates as assessed by CALV1 and CALV12. Increased proportion of NAHF genes exhibited a negative effect on survival (P<0·001) whereas increased levels of heterosis had a positive impact on survival (P<0·001). The results of the present study indicate that in seasonal calving herds in Ireland a need for direct selection on traits related to fertility and survival is required to arrest and reverse the declining trends in calving rates and survival.

Effects of calving age, breed fraction and month of calving on calving interval and survival across parities in Irish spring-calving dairy cows

The objective of this study was to investigate the effect of parity, age at calving, percentage North American Holstein-Friesian and calving date on subsequent calving interval and survival to facilitate the estimation of transition probabilities for month of calving. The economic value of traits that influence calving date, age distribution and survival can be assessed in models using a transition probability matrix. Such a matrix contains the probabilities that a cow of a particular age or breed calving in a particular month will calve in the same, an earlier or later month next year, or be culled. Following editing 1,046,855 calving records in spring-calving herds between the years 1990 and 2004 were analysed. Shorter calving intervals were associated with cows calving later in the calendar year. Age at first calving of < 24 months resulted in longer calving intervals to second calving across all levels of Holstein percentage with cows calving for the first time at 25–26 months of age having the shortest subsequent calving interval. Age at second calving of 37–38 months and third calving of 49–50 months were optimum for shorter subsequent calving intervals. Calving interval increased with Holstein percentage across the first 5 parities. Survival rate decreased with later month of calving and with older parities. When survival rate was measured as the ability of the cow to re-calve within 500 days, the highest survival rate was found in cows calving at 25–26 months of age whereas there was a noticeable reduction in survival across all parities in the 88–100% Holstein percent category.

Consequences of genetic selection for increased milk production in European seasonal pasture based systems of milk production

The data presented in this review show that past selection for increased milk production in Ireland over a 14-year period (1990 to 2003) resulted in increases in milk production per cow but has been accompanied by undesirable side effects in reproduction and survival. Economic analysis in a EU milk quota scenario over this period showed that only 41% of the potential improvement in farm profit was achieved because of impaired reproductive performance. Strain comparison studies show that genetic selection for increased phenotypic milk production, which is generally accompanied with increased proportion of North American Holstein Friesian genes, was associated with increased milk production per cow, lower body condition score, greater milk production response to concentrate supplementation at pasture and reduced fertility and survival. The lower reproductive performance observed with cows selected solely for high milk production was associated with lower energy balance in early lactation, greater partitioning of additional nutrients towards milk production, inability to achieve desired grass intake from pasture and decreased plasma glucose and insulin like growth factor-1 concentrations. The existence of genotype by environment interactions suggests that germplasm selected in the future should be from within the management system in which it is to be used, and based on a selection index that combines production and other traits of economic importance.

Nutritional limitations to increased production on pasture-based systems.

The constraints to high levels of milk production imposed by a high-quality-pasture diet, and development of feeding strategies to overcome these limitations, were examined by modelling the nutritional status of New Zealand Friesian and North American Holstein-Friesian dairy cows grazing high-quality pasture. The Cornell Net Carbohydrate and Protein System (CNCPS) was used to predict sensitivity of milk production to a 10% change in the composition of pasture nutrients. The rate at which fibre and protein were degraded in the rumen and the value given to effective fibre and lignin content significantly affected the supply of metabolisable energy and protein, and the profile of amino acid supply. The first limiting factor in milk production when only high-quality pasture was fed was metabolisable energy supply, while specific amino acids, particularly methionine and lysine, limited milk production when > 20 g/kg diet consisted of a grain supplement. Compared with cows fed a total mixed ration in confinement, North American Holstein-Friesians grazing all pasture produced less milk (29.6 v. 44.1 kg/d). Of the difference in milk production 61% could be attributed to a lower DM intake (19 kg/d v. 23.4 kg/d). Predictions using the CNCPS indicated that supply of metabolisable energy was the first-limiting factor for milk production from high-quality pasture (251 g crude protein (N x 6.25)/kg, 432 g neutral-detergent fibre/kg, 77% in vitro DM digestibility), rather than metabolisable protein or amino acids. In addition, these nutritional limitations imposed by pasture diets will be greater for dairy cow genotypes that have not been selected for high performance within a pasture system.

Is it feasible to use ovum pick up and embryo transfer to produce 3 bull calves per donor and maintain annual calving cycles?

The objective of this study was to evaluate the effect of stocking rate (SR) and calving date (CD) on milk production, BW and BCS within grass-based production systems post-European Union milk quotas using modern grazing management practices and high genetic potential Holstein–Friesian (HF) spring calving dairy cattle over a two year period. Two groups of HF dairy cows with different mean CD were established from within the existing research herd at Moorepark (Teagasc, Ireland). Animals were assigned either to an early calving (mean CD: 12 February) treatment or a late calving (mean CD: 25 February) treatment. Animals within each CD treatment were randomly allocated to 1 of 3 SR treatments, Low (2.51 cows/hectare (ha), Medium (2.92 cows/ha) and High (3.28 cows/ha) which were designed to represent 3 alternative whole farm SR in a post-European Union milk quota, spring calving, grass-based milk production system. A total of 138 spring-calving dairy cows, comprised of two strains of HF (North American HF and New Zealand HF genetic strains), were used during 2009 and 2010, respectively. The effects of CD, SR treatment, genetic strain and their interactions on milk production per cow and per ha, body weight and body condition score were analyzed. Although reducing per animal production, increased SR resulted in increased milk and milk solids production per ha. The results also indicate that although CD had no effect on total lactation performance, adjusting mean CD may be an effective strategy to align animal requirements and grass supply and reduce the requirement for supplements at increased SR in early lactation.