Moreno Formation Research Articles

Elasmosaurid phylogeny and paleobiogeography, with a reappraisal of Aphrosaurus furlongi from the Maastrichtian of the Moreno Formation

ABSTRACTThe holotype of the elasmosaurid Aphrosaurus furlongi from the Maastrichtian levels of the Moreno Formation is redescribed and considered a valid species based on one autapomorphy, a deep trough in the ventral surface of vertebral centra of the posterior cervicals, and the following combination of features: wide and short clavicle-interclavicle complex with concave anterior border and without posterior medial process or medial ventral keel, humerus with anterior depression, and posterior limb with accessory element on the posterior margin. The phylogenetic analysis recovered A. furlongi as an elasmosaurid within Weddellonectia. The phylogenetic analysis generated a new topology of Elasmosauridae. A new clade, Euelasmosaurida, is recovered, including all post-Cenomanian elasmosaurids other than Zarafasaura oceanis. Euelasmosaurida is composed of two main clades: Elasmosaurinae and Weddellonectia. The latter includes mostly Weddellian elasmosaurids, including Aristonectinae. Based on the new phylogenetic results, the evolution of key cranial and postcranial characters is discussed. Two key intervals of elasmosaurid evolutionary history are recognized: the Cenomanian, with the appearance of Euelasmosaurida, and the Santonian, with the differentiation of Weddellonectia and Elasmosaurinae.

Mechanisms of biogenic gas migration revealed by seep carbonate paragenesis, Panoche Hills, California

A comprehensive study of seep carbonates at the top of the organic- rich Maastrichtian to Danian Moreno Formation in the Panoche Hills (CA) reveals the mechanisms of generation, expulsion and migration of biogenic methane which fed the seeps. Two selected outcrops show seep carbonates developed at the tip of sand dykes intrude up into the Moreno Formation from deeper sandbodies. Precipitation of methane-derived cements occurred in a succession of up to 10 repeated elementary sequences, each starting with a corrosion surface followed by dendritic carbonates, botryoidal aragonite, aragonite fans and finally laminated micrite. Each element of the sequence reflects three stages. First a sudden methane pulse extended up into the oxic zone of the sediments, leading to an aerobic oxidation of methane and carbonate dissolution. Secondly, after consumption of the oxygen, anaerobic oxidation of methane coupled with sulfate reduction triggered carbonate precipitation. Third, progressive diminishment of the methane seepage, lead to the deepening of the reaction front in the sediment and a lowering of precipitation rates. Carbonate isotopes, with δ13C as low as -51‰PDB, indicate a biogenic origin for the methane, while a 1D basin model suggests that the Moreno Formation was in optimal thermal conditions for bacterial methane generation at the time of seep carbonate precipitation. Methane pulses are interpreted to reflect drainage by successive episodes of sand injection into the gas-generating shale of the Moreno Formation. The seep carbonates of the Panoche Hills can thus be viewed as a record of methane production from a biogenic source rock by multiphase hydraulic fracturing.

Reappraisal of Fresnosaurus drescheri (Plesiosauria; Elasmosauridae) from the Maastrichtian Moreno Formation, California, USA

The elasmosaurid Fresnosaurus drescheri, Welles from the contact between the Tierra Loma/Marca members of the Moreno Formation (Maastrichtian), California, USA is reviewed. Most of the features included in Welles's original diagnosis are considered related only to the juvenile ontogenetic stage of the holotype and only specimen. The new diagnosis is based on diagnostic characters of the ilia, including long rectangular shaped sacral facets located in the dorsal part of the shaft, two gentle knobs in the shafts and unexpanded dorsal end. Additional material from the Moreno Formation (numbered under the same number as the F. drescheri holotype but not mentioned by Welles and therefore considered part of a different specimen) are described for the first time. The latter are referred to the aristonectine, being the first evidence of aristonectines from the North Hemisphere.

The late-surviving ‘duck-billed’ dinosaur Augustynolophus from the upper Maastrichtian of western North America and crest evolution in Saurolophini

AbstractWe amend the taxonomy and provide new anatomical information on the hadrosaurid dinosaur Saurolophus morrisi (upper Maastrichtian Moreno Formation, central California, USA) derived from full preparation of the referred skull roof. The cranial morphology of this species is distinct enough to justify the new combination Augustynolophus morrisi gen. nov. The morphology of the nasals and surrounding cranial bones indicates that A. morrisi sported a solid nasal crest ending in an elongate triangular plate that extended above the skull roof. Autapomorphies include a crescentic base of the frontal caudodorsal process and extension of the process caudal to the frontal ‘dome’; distal end of nasal crest with knob-like process inflected rostrally; circumnarial depression lightly incised and weakly emarginated, adjacent to caudolateral margin of nasal and occupying two-thirds the width of lateral surface of distal region of crest; and caudal surface of distal nasal crest subrectangular. We formally establish the new tribe Saurolophini consisting of Prosaurolophus, Augustynolophus and Saurolophus. Saurolophin synapomorphies include a premaxilla with broad arcuate contour of rostrolateral region of thin everted oral margin and flat and steeply inclined occlusal surface of dentary dental battery, among other characters. Saurolophin crests evolved towards increasing caudodorsal length, along with caudal extension of the circumnarial fossa and involvement into the crest of adjacent facial elements. Augustynolophus is the second described genus of North American late Maastrichtian hadrosaurids. Its recognition implies a greater diversity among late Maastrichtian dinosaur faunas than previously recognized and is congruent with hypotheses of endemism and/or provinciality during Late Cretaceous time.

A basin modeling and organic geochemistry study in the Vallecitos syncline, San Joaquin Basin, California

The Vallecitos syncline is a westerly structural extension of the San Joaquin Basin. The Vallecitos oil field, comprised of eight separate areas that produce from Cretaceous and Paleogene reservoirs, accounted for 5.4 MMB of oil and 5.6 BCF associated of gas through 2010. However, exploration for oil and gas in the Vallecitos area is challenging due to structural complexity and limited data. The purpose of this study is to evaluate whether source rocks are actively generating petroleum in the Vallecitos syncline and to improve our understanding of burial history and timing of hydrocarbon generation. We conducted biomarker analysis on twenty-two oil samples from the Vallecitos syncline. Source-related biomarkers show two genetic groups of oil, which originated from two different source rocks. These results differ from earlier published interpretations in which the Kreyenhagen Formation is the only source rock in the Vallecitos syncline, and suggest that the Cretaceous Moreno Formation in the syncline also is an active source rock.Stratigraphic evidence and modeling suggest that late Cenozoic episodes of erosion due to folding and uplift removed significant overburden on the flanks of the syncline. To better understand the petroleum systems and clarify the total active source rocks in the area, 2D burial histories were generated through the Vallecitos syncline. A published cross-section through the deepest part of the syncline was selected to conduct thermal history, basin evolution, and migration analyses. The 2D model results indicate that the lower Kreyenhagen Formation has various maturities within the formation at different locations in the present-day syncline. The basal part of the Kreyenhagen Formation is in the dry gas window and maturity decreases away from the central part to the flanks. It remains immature along shallow portions of the present-day flanks. In contrast, the basal part of the Moreno Formation achieved extremely high maturity (past the gas generation zone) but is in the oil generation zone on the flanks of the syncline at shallow depth. All of our geochemical and 2D model results suggest that there are two active source rocks in the Vallecitos syncline. Accordingly, we propose that there are two active petroleum systems in the Vallecitos syncline.

Aquatic adaptation, cranial kinesis, and the skull of the mosasaurine mosasaur Plotosaurus bennisoni

ABSTRACT New anatomical observations of the holotype skull of Plotosaurus bennisoni from the Maastrichtian Moreno Formation of California, U.S.A., are used as a framework to examine cranial kinesis in derived members of the Mosasaurinae. Enlarged posteromedial flanges of the frontal and extensive lateral contacts of the prefrontal and postorbitofrontal contributed to increased rigidity along the frontoparietal suture (the mesokinetic joint). Sutural contacts of the parietal with the supraoccipital posteriorly and the prootic ventrally would have restricted metakinetic movements. Furthermore, the unusual shape of the epipterygoid, and its dorsal contact with the prootic and parietal, shows that the epipterygoid and pterygoid were probably not capable of anteroposterior movements. Most strikingly, Plotosaurus exhibits a tight association of the quadrate with the temporal arcade, suggesting that streptostyly was limited or lost in this derived mosasaurine, the loss of such a feature having never been described in a mosasaur. These charcteristics are placed in a functional context to examine aquatic adaptations in mosasaurs. As one of the most specialized mosasaurs known, the loss of cranial kinesis may have evolved as a result of its piscivorous diet.

Geographic and stratigraphic distribution of the Late Cretaceous suspension-feeding bony fish Bonnerichthys gladius (Teleostei, Pachycormiformes)

ABSTRACT The stratigraphic and paleogeographic distribution of the suspension-feeding pachycormiform fish Bonnerichthys is reviewed. Fossils attributable to this genus are known from the Western Interior Seaway (Niobrara Formation of Kansas, Sharon Springs Formation of Kansas, Nebraska, and North Dakota, DeGrey Formation of South Dakota, and possibly Mobridge Formation of Nebraska), the Eastern (Eutaw Formation of Mississippi and Mooreville and Demopolis formations of Alabama and Mississippi) and Western (Ozan Formation of Texas and Marlbrook Marl Formation of Arkansas) Gulf Coastal Plain, the Atlantic Coastal Plain (Wenonah Formation of New Jersey and possibly Marshalltown Formation of Delaware), and Pacific coast (Moreno Formation of California). Chronologically, occurrences of Bonnerichthys range in age from Coniacian to Maastrichtian. Fossils of Bonnerichthys from the Atlantic Coastal Plain, Eastern and Western Gulf Coastal Plain, and Pacific coast are definitively identified and figured here for the first time. Candidate deposits outside the United States that might yield Bonnerichthys are reviewed.

Description of new specimens of Halisaurus arambourgi Bardet & Pereda Suberbiola, 2005 and the relationships of Halisaurinae

Abstract Halisaurine mosasaurs are poorly known, represented by a small number of specimens from the Santonian-Maastrichtian (~86 Ma – ~66 Ma), but enjoyed broad palaeobiogeographic distribution during that time. They are important for understanding mosasaur evolution because certain aspects of their morphology retain the relatively plesiomorphic or minimally modified squamate conditions; however, existing material is limited and certain anatomical details are lacking. We report here two new specimens of Halisaurus arambourgi including a well-preserved, nearly complete skull and postcranial skeleton, and a partial skull that preserves details of the braincase and quadrate. We focus our description on morphology that augments the original description of this species and provides comparisons with other halisaurines. Braincase and temporal arcade characters confirm the plesiomorphic nature of Halisaurus, supporting a relatively basal position of Halisaurinae within Mosasauridae. Comparisons of cranial morphology support reconstruction of relationships within Halisaurinae, indicating that H. arambourgi is most closely related to H. platyspondylus, Phosphorosaurus (= H. ortliebi) is the sister taxon to those taxa, and Eonatator is the most basal described halisaurine. The proportions of the epipodials and the caudal vertebral centrum morphometrics indicate H. arambourgi is more derived than the Santonian to early Campanian Eonatator sternbergii but less derived than a Halisaurus sp. specimen from the mid-Maastrichtian of the Moreno Formation of California, USA. Moreover, vertebral morphometrics reveals that H. arambourgi possessed a downturned tail that likely supported a crescent-like fluke.

Saurolophus morrisi, a new species of hadrosaurid dinosaur from the Late Cretaceous of the Pacific coast of North America

We describe and re-evaluate the systematics of specimens from the Maastrichtian Moreno Formation of California (western USA) as a new species of Saurolophus, the only known genus of hadrosaurid dinosaur widespread in Asia and North America. Recognition of this new species adds substantially to the record of the taxonomic diversity of these animals west of the Rocky Mountains. The new species, Saurolophus morrisi, is diagnosed by the possession of a postorbital having ornamentation in form of wide oblique groove on jugal process. Placement of this new species in Saurolophus considerably expands the distribution of this genus, although this referral is arbitrary since phylogenetic analysis places the new species outside of the clade formed by Saurolophus osborni and Saurolophus angustirostris. However, recognition of a new, endemic Californian hadrosaurid, especially one so closely related to both Asian and North American species, may have implications for future studies of both the internal biogeography of...

Revision of the status of Saurolophus (Hadrosauridae) from California, USA

The occurrence of Saurolophus from the Moreno Formation (late Maastrichtian) of California is investigated and an incomplete, poorly preserved, skull (LACM/CIT 2852) is described. The skull lacks the braincase (including the frontals) and much of the nasals, and the preserved portions are crushed or plastically deformed, which makes anatomical interpretations difficult. A preserved midline fragment of the conjoined nasals suggests that it lacked a gryposaur-like “Roman nose”, but the nature of the crest, if present, is impossible to determine with certainty. A phylogenetic analysis places this specimen as either the sister taxon of Saurolophus or as the sister taxon to a clade comprising Edmontosaurus and Anatotitan . There is no compelling morphological evidence to support the previous assignment of LACM/CIT 2852 to Saurolophus rather than to Edmontosaurus, and its poor preservation prevents positive assignment to any taxon below Hadrosaurinae indet. Given its geographic setting and morphological uncertainties, it is also possible that this specimen represents a separate taxon, but more material is needed to clarify the identity of the Moreno hadrosaurine. LACM/CIT 2852 does, however, provide evidence that Maastrichtian hadrosaurines ranged west of the Sierra Nevada magmatic arc, in an area where dinosaur diversity is poorly known.

Biostratigraphic and biogeographic implications of a hadrosaurid (Ornithopoda: Dinosauria) from the Upper Cretaceous Almond Formation of Wyoming, USA

The results of Barnum Brown's 1937 expedition to the Almond Formation of Wyoming consisted of two unidentified ceratopsian skulls and a partial hadrosaurid specimen (AMNH 3651). The hadrosaurid is here attributed to the Maastrichtian genus Saurolophus, verifying previous biostratigraphic correlations of this formation using ammonite zones. Fossiliferous lower Maastrichtian formations occurring latitudinally between those of Alberta, Canada, and southwestern Texas, USA, such as the Almond Formation, are essential for testing the effects and duration of apparent hadrosaurid faunal segregation earlier in the Campanian, and indirectly aiding in the placement of faunal boundaries that are currently unknown for the late Campanian. The discovery of Saurolophus in Wyoming, a close relative of the Campanian genus Prosaurolophus, affirms that the segregation of hadrosaurid faunas established in the late Campanian (∼75 Ma) continued for at least 3 million years. Combining occurrences of Saurolophus from Mongolia and the Moreno Formation of California with those of Alberta, Canada, this genus appears to have had one of the largest geographic ranges of any equivalent clade of hadrosaurid dinosaur, although species level distributions are still uncertain.

Cranial osteology of the giant mosasaurPlesiotylosaurus(Squamata, Mosasauridae)

Based on the type material and a new, virtually intact skull from the mid-Maastrichtian part (probably Tierra Loma Shale) of the Moreno Formation, San Joaquin Valley, central California, the cranial osteology of the enigmatic mosasaurinePlesiotylosaurus crassidens(Squamata, Mosasauridae) is here described in detail.Plesiotylosaurusis characterized by a protruding rostrum on the snout and mandibles, a wide and virtually unconstricted internarial bar, large and robust pterygoid teeth, and quadrates with distally fused suprastapedial and infrastapedial processes. Distinctly obtruding predental prows, along with a number of elaborate cranial features, suggest a morphological convergence with tylosaurine mosasaurs.

An ancient linked fluid migration system: cold-seep deposits and sandstone intrusions in the Panoche Hills, California, USA

In central California, Maastrichtian–Danian shales of the Moreno Formation preserve a fluid migration system that developed along the western margin of the former Great Valley forearc basin. The system consists of a network of interconnected sandstone intrusions linked to overlying fossiliferous carbonates whose geochemistry, fauna, and petrology are characteristic of active cold seeps. The system is approximately 800 m thick and represents episodic migration and seafloor expulsion of fluids over at least 0.5×106 years. This locality has the most extensive exposure yet discovered of a complete seep system, from underlying fluid pathways to seep deposits and associated communities.

NEW LATE CRETACEOUS (CAMPANIAN AND MAASTRICHTIAN) MARINE GASTROPODS FROM CALIFORNIA

Three new genera and six new species of shallow-marine gastropods are named from Upper Cretaceous strata found mainly in California. The trochids Cidarina cretacea new species and Cidarina beta new species, the ficid Bulbificopsis garza new genus and new species, and the cancellariid Mataxa arida new species are from the Maastrichtian part of the Moreno Formation of north-central California. This is the earliest record of Cidarina, whose previous chronologic range was middle Eocene to Recent. Bulbificopsis is the first record of a Cretaceous ficid from the Pacific slope of North America, and Mataxa was previously known only from Upper Cretaceous strata in the southeastern United States and northeastern Brazil. The buccinid Eripachya jalama new species and the fasciolariid Calkota daileyi new genus and new species are from the lower upper Campanian Jalama Formation in southern California. Calkota is also recognized herein as occurring in upper Maastrichtian strata of North Dakota and South Dakota. The new melongenid genus, Pentzia, established for Fulgur hilgardi White, 1889, is from Campanian strata throughout California; middle Campanian strata on Sucia Island, Washington; and upper Campanian to lower Maastrichtian strata in northern Baja California, Mexico.

New late Cretaceous (Campanian and Maastrichtian) marine gastropods from California

Three new genera and six new species of shallow-marine gastropods are named from Upper Cretaceous strata found mainly in California. The trochidsCidarina cretaceanew species andCidarina betanew species, the ficidBulbificopsis garzanew genus and new species, and the cancellariidMataxa aridanew species are from the Maastrichtian part of the Moreno Formation of north-central California. This is the earliest record ofCidarina, whose previous chronologic range was middle Eocene to Recent.Bulbificopsisis the first record of a Cretaceous ficid from the Pacific slope of North America, andMataxawas previously known only from Upper Cretaceous strata in the southeastern United States and northeastern Brazil. The buccinidEripachya jalamanew species and the fasciolariidCalkota daileyinew genus and new species are from the lower upper Campanian Jalama Formation in southern California.Calkotais also recognized herein as occurring in upper Maastrichtian strata of North Dakota and South Dakota. The new melongenid genus,Pentzia, established forFulgur hilgardiWhite, 1889, is from Campanian strata throughout California; middle Campanian strata on Sucia Island, Washington; and upper Campanian to lower Maastrichtian strata in northern Baja California, Mexico.

Marine reptiles from the Nanaimo Group (Upper Cretaceous) of Vancouver Island

A new fauna of fossil marine reptiles is described from the Late Cretaceous Nanaimo Group of Vancouver Island. The fossils are from the Haslam and Pender formations (upper Santonian) near Courtenay, British Columbia, and include elasmosaurid plesiosaurs, turtles, and mosasaurs. This is only the second fauna of Late Cretaceous marine reptiles known from the Pacific Coast, the other being from the Moreno Formation of California (Maastrichtian). The new Nanaimo Group fossils are some 15 million years older than those from the Moreno Formation. However, like the California fauna, there are no polycotylid plesiosaurs, and one of the mosasaurs is a new genus. This reinforces the provinciality of the Pacific faunas and their isolation from contemporaneous faunas in the Western Interior Seaway.

Cyclic Organic-Rich Sedimentation in the Marca Shale, Moreno Formation and the Onset of Late Cretaceous Upwelling along the California Margin: ABSTRACT

Cyclic Organic-Rich Sedimentation in the Marca Shale, Moreno Formation and the Onset of Late Cretaceous Upwelling along the California Margin: ABSTRACT

70 Million Years of Coastal Upwelling in California; Source Rocks and Paleoceanography: ABSTRACT

The Late Mesozoic-Cenozoic marine sequence of California displays a unique record of coastal upwelling and productivity in the form of distinctive diatomaceous and organic-rich deposits including the upper Cretaceous-lower Paleocene Moreno Formation, the Eocene Kreyenhagen Formation and the Miocene Monterey Formation. Unique sedimentological characteristics of these ancient upwelling systems include (a) Finely laminated biosiliceous-rich shales (>30% biogenic silica content), (b) Distinctive laminae composed by monospecific diatom resting spores, (c) Good source rock quality (>300 mg HC/mg org C), and (d) High accumulation rates in mid water anoxic conditions. Detailed study of individual laminae in sediments of these formations revealed concentration of monospecific diatom resting spores and an abundance of Stephanopyxis sp. and Coscinodiscus sp. Like Recent upwelling systems, preserved laminations of monospecific resting spores reflect strong seasonal upwelling that lead to high organic matter production and enhancement of anoxia. The presence of spores in the ancient and modern upwelling systems is evidence that diatoms have adapted and successfully competed in the highly productive California Margin since the Late Cretaceous. The Moreno, the Kreyenhagen and the Monterey Formation account for a significant portion of major petroleum source beds in California and contain an important record of coastal upwelling and paleoceanographic changemore » in the northeastern Pacific Ocean over the past 70 million years. It is suggested that potential Late Maestrichtian source rocks could have been deposited along other favorable upwelling areas of the Eastern Pacific Rim.« less

Geochemical evidence for Se mobilization by the weathering of pyritic shale, San Joaquin Valley, California, U.S.A.

Acidic (pH 4) seeps issue from the weathered Upper Cretaceous-Paleocene marine sedimentary shales of the Moreno Formation in the semi-arid Coast Ranges of California. The chemistry of the acidic solutions is believed to be evidence of current reactions ultimately yielding hydrous sodium and magnesium sulfate salts, e.g. mirabilite and bloedite, from the oxidation of primary pyrite. The selenate form of Se is concentrated in these soluble salts, which act as temporary geological sinks. Theoretically, the open lattice structures of these hydrous minerals could incorporate the selenate (SeO 4 −2) anion in the sulfate (SO 4 −2) space. When coupled with a semi-arid to arid climate, fractional crystallization and evaporative concentration can occur creating a sodium-sulfate fluid that exceeds the U.S. Environmental Protection Agency limit of 1000 μg l −1 for a toxic Se waste. The oxidative alkaline conditions necessary to ensure the concentration of soluble selenate are provided in the accompanying marine sandstones of the Panoche and Lodo Formations and the eugeosynclinal Franciscan assemblage. Runoff and extensive mass wasting in the area reflect these processes and provide the mechanisms which transport Se to the farmlands of the west-central San Joaquin Valley. Subsurface drainage from these soils consequently transports Se to refuge areas in amounts elevated to cause a threat to wildlife.

Stratigraphic Variations in the Biomarker Distribution of the Moreno Formation: Their Correlation with San Joaquin Basin Oils: ABSTRACT

Stratigraphic Variations in the Biomarker Distribution of the Moreno Formation: Their Correlation with San Joaquin Basin Oils: ABSTRACT