Tournefortia gigantifolia, an unusual understory shrub, is described as new from wet forests in Ecuador and Colombia. It is distinct in its monocaulous or sparsely branched habit with large punctate leaves clustered near the stem apices. Its free anthers and unlobed fruits indicate clearly that it is a member of Tournefortia sect. Tournefortia, but its relationships with other species are not evident. RESUMEN. Tournefortia gigantifolia, un arbusto excepcional de sotobosque, se describe como nueva especie. Habita en bosques htimedos en Ecuador y Colombia. Es distintivo en su forma de crecimiento monocaule o con pocas ramas, con hojas grandes que presentan abundantes puntos en el haz; las hojas estan agrupadas cerca de los Apices de los tallos. Aunque esta nueva especie claramente pertenece al g6nero Tournefortia, secci6n Tournefortia, sur parentesco con las otras especies de la secci6n no esti claro. The pantropical genus Tournefortia L. comprises at least 150 species that are strongly centered in the Neotropics. Most modem authors have treated the genus in a broad sense (Nowicke, 1969; Gibson, 1970; Nowicke & Skvarla, 1974) and have included species in three sections, although the nomenclature has been confused. Johnston (1930) recognized section Cyphocyema I. M. Johnston, a group of perhaps 20 species confined to the New World tropics, and section Tournefortia (as sect. Eutournefortia), which comprises the vast majority of the genus with well over 130 species. One species of section Tournefortia occurs in eastern Africa (Verdcourt, 1988), four are found in Madagascar and the Mascarenes, and eight range from southern Asia through the Philippines to Australia (Johnston, 1935b). The more than 100 species that occur in tropical America are mostly South American, and the majority of these species are found in the Andean countries. The third section, sect. Mallatonia Grisebach, consists of three species widely distributed in the Caribbean, Indian, and Pacific oceans. Johnston's opinion on the placement of these species varied widely through his publications on the gen s (Johnston, 1930, 1935a, b, 1949, 1951), and some authors have followed Britton (1915) in recognizing these species as generically distinct, but they have often been grouped together. The genus has all but been ignored by modem monographers, and the last comprehensive treatment was that of De Candolle (1845). Since that time, the New World species have been treated only in floristic works and regional studies. The Central American species are comparatively well studied, having been treated for Veracruz (Nash & Moreno, 1981), Guatemala (Gibson, 1970), Nicaragua (Miller, in press), and Panama (Nowicke, 1969; Miller, 1988). Two studies of South American species of Tournefortia were published by Ivan Johnston that treated the species of Brazil, Paraguay, Uruguay, and Argentina (Johnston, 1930) and the Guianas and adjacent Brazil north of the Amazon (Johnston, 1935a). Unfortunately, the Andean countries, certainly the center of diversity for the genus, have largely been ignored. E. P. Killip prepared, but never published (see Nowicke, 1974, for a more complete discussion of Killip's unpublished manuscripts), a treatment for the Andean countries, including Venezuela, Colombia, Ecuador, Peru, and Bolivia, and Killip's manuscript still exists in the archives of the National Herbarium at the Smithsonian Institution. While never submitted for publication, Killip's work did serve as the basis for two papers describing new species from the region (Killip, 1927, 1929). One more of the species recognized as new by Killip was later published by Nowicke (1974). Since Killip's effort there has been no comprehensive study of the Andean species, and new taxa from the northern Andean countries have been described individually (Johnston, 1956; Nowicke, 1974; Gentry, 1977; Gilli, 1983; Miller, 1989, 1995, 1997, 2000). While Killip's short papers published most of the new taxa that he recognized, several were never validly published; their use has nevertheless crept into herbaria from specimens that Killip annotated. One of these is for a plant from wet forests of northern Ecuador and adjacent Colombia that has never been described despite NovoN 10: 48-52. 2000. This content downloaded from 157.55.39.132 on Thu, 15 Sep 2016 05:51:44 UTC All use subject to http://about.jstor.org/terms