Mature Wood Zones Research Articles

LATERAL GROWTH AFFECTS XYLEM MATURATION PROPERTIES OF 8-YEAR-OLD BIG-LEAF MAHOGANY (SWIETENIA MACROPHYLLA)

The area of juvenile wood zone of an 8-year-old planted big-leaf mahogany (Swietenia macrophylla) was uniform regardless of diameter. Hence, it was assumed that xylem maturation is dependent on diameter. Radial distribution patterns with reference from the pith exhibited short fibre and vessel lengths, narrow vessel width and low xylem density, and exhibited gradual increase outward and became more or less stable near the bark. However, the radial distribution patterns of fibre width and modulus of elasticity were scattered regardless of tree diameter sizes. Fibre length, vessel length, vessel width and xylem density of the juvenile wood zones showed significant differences compared with the transition and mature wood zones. However, no significant difference was observed between transition wood zone and mature wood zone except for fibre length. A negative correlation was observed in fibre length, vessel length, vessel width and xylem density in terms of their relationship with diameter at breast height and the radial distribution of fibre (i.e. b-value), and the diameter boundaries of the three wood zones. At diameters 18.08, 17.36, 16.23 and 17.87 cm respectively, fibre length, vessel length, vessel width and xylem density can be used to identify xylem maturation boundary.

Interpreting research results for the physical and mechanical properties of wood: An approach not dependent on a juvenile/mature wood boundary

The interpretation and presentation of research on the physical and mechanical properties of wood in the radial direction is important for the estimation of technological properties in primary wood processing. It is common practice to define the boundary between the juvenile and mature wood zone of tree growth because of the differences in wood properties in these two zones. The juvenile and mature wood zones can be determined statistically based on the significance of the difference in the properties in a particular zone. This paper presents the insufficiency in the statistical determination of the boundary between juvenile and adult wood. Such limitations detract from the potential value and technological exploitation of wood as raw material. Statistical tests yielded zones that were too wide for the transition of juvenile wood to mature wood. Representations of the distribution of properties in the radial direction also complement the knowledge for assessing the technological properties based on the researched use of the presentation of polynomials of the second degree and the display of the Tukey HSD test in the form of comparison tables. The graphical representations by groups of the tested annual rings of fir wood also help to assess the technological properties.

Radial variation in tracheid lengths in dominant trees of selected coniferous species

The radial variation was examined for tracheid lengths of Norway spruce (Picea abies (L.) Karst.), European larch (Larix decidua Mill.), and Scots pine (Pinus sylvestris L.) wood from dominant trees coming from an even-aged stand, and growing under identical forest site and climatic conditions. The measurements were completed on macerated material. The variation of tracheid lengths in annual rings from the core to the bark was used for determination of the border between the juvenile and mature wood in the trunk cross-section. The boundary age between the juvenile and mature wood zones established for the examined species was comparable, as it was 25 annual rings for Scots pine and 29 for European larch and for Norway spruce. In the juvenile zone, the tracheid lengths increased 2.2-fold in Norway spruce wood, while in Scots pine and European larch wood it was approximately 1.7-fold. By contrast, in the mature wood zone the tracheid lengths was stabilized at a certain level, showing slight fluctuations. The differences in the tracheids length of early and late wood in the examined annual rings were also determined, and it was established that for the majority of annual rings they are statistically significant (p<0.05).

Wood properties of Carapa guianensis from floodplain and upland forests in Eastern Amazonia, Brazil

The variable environmental conditions of the Amazon forest can affect the wood properties of the tree species distributed across its diverse phytophysiognomies. Carapa guianensis (Andiroba) occurs in upland and floodplain forests, and the wood and oil of its seeds have multiple uses. The aim of this study was to evaluate the wood properties of C. guianensis trees in upland and estuarine floodplain forests of the Amazon River. Eight trees were selected, with four being from the upland and four from the floodplain forests. The fiber length, fiber wall thickness, vessel diameter and frequency, microfibril angle, specific gravity and wood shrinkage were evaluated. The juvenile and mature wood zones were determined according to these variables. The fiber length, fiber wall thickness and specific gravity increased, and microfibril angle decreased, in the pith to bark direction. Only the fiber length variable was efficient for delimiting juvenile, transition and mature wood. The fiber length, wall thickness and specific gravity of wood were higher in upland forest trees. However, the environment did not alter the beginning of the formation and proportion of C. guianensis mature wood. This information is important for the log fit in cutting diagrams, aiming toward improving the production, classification and processing of pieces with specific quality indexes in order to direct them to appropriate wood uses.

Phenotypic Correlations among Growth and Selected Wood Properties in White Spruce (Picea glauca (Moench) Voss) †

We examined phenotypic relationships among radial growth-related, physical (i.e., related to wood density), and anatomical (i.e., related to tracheid dimensions) wood properties in white spruce (Picea glauca (Moench) Voss), in order to determine the strength and significance of their correlations. Additionally, principal component analysis (PCA) was used to establish if all of the properties must be measured and to determine the key properties that can be used as proxies for the other variables. Radial growth-related and physical properties were measured with an X-ray densitometer, while anatomical properties were measured with a Fiber Quality Analyzer. Fifteen wood properties (tracheid length (TL) and diameter (TD), earlywood tracheid length (ETL) and diameter (ETD), latewood tracheid length (LTL) and diameter (LTD), ring width (RW), ring area (RA), earlywood width (EWW), latewood width (LWW), latewood proportion (LWP), ring density (RD), intra-ring density variation, earlywood density (EWD), and latewood density (LWD)) were assessed. Relationships were evaluated at intra-ring and inter-ring levels in the juvenile wood (JW) and mature wood (MW) zones. Except for a few cases when mature tracheid diameter (TD) was involved, all intra-ring anatomical properties were highly and significantly correlated. Radial growth properties were correlated, with stronger relationships in MW compared to JW. Physical properties were often positively and significantly correlated in both JW and MW. A higher earlywood density coupled with a lower latewood density favored wood uniformity, i.e., the homogeneity of ring density within a growth ring. Managing plantations to suppress trees growth during JW formation, and enhancing radial growth when MW formation starts will favor overall wood quality. In order, RW-EWW-RA, TL-ETL-LTL, and RD-EWD-LWP are the three clusters that appeared in the three wood zones, the whole pith-to-bark radial section, the juvenile wood zone, and the mature wood zone.

Physical Properties of Juvenile and Mature Sycamore Maple (Acer pseudoplatanus L.) Wood from Medvednica Region

Physical properties play an important role in predicting wood quality. The aim of this study was to investigate radial variations in physical properties of sycamore maple (Acer pseudoplatanus L.) wood, to determine the border between juvenile and mature wood and to compare them with analogous variations in beech wood. Radial variation of density in absolutely dry condition, maximum density, nominal density, longitudinal, radial, tangential and volumetric shrinkage, maximum moisture content and annual ring width on five sycamore maple trees from Medvednica region in Croatia were determined. Observing the trend curves and comparing average values of physical properties, different zones in the cross section of the trunk were determined. In the first zone, from the pith to about 30th annual ring, more rapid changes in physical properties occurred. It is followed by transitional zone, which ranges roughly from 30th to 40th annual ring. After 40th annual ring, less pronounced changes in measured physical properties occurred. Sharp boundary between juvenile and mature wood zones could not be determined. According to the results, the first zone is considered to be juvenile wood, while mature wood starts after the 30th annual ring .

Wood Variability in Mature Longleaf Pine: Differences Related to Cardinal Direction for a Softwood in a Humid Subtropical Climate

Mature longleaf pine ( Pinus palustris Mill .) trees were harvested to compare wood property data for opposing bark-to-pith wood strips representing the northern and southern cardinal (or compass) directions. For each of the ten 70-yr-old trees used in the study,wood property datawere compared at breast height (BH) and three relative heights: one-quarter height (1Q), midheight (MID), and three-quarter height (3Q). Scanning of the specimens by X-ray densitometry gave specific gravity (SG) profiles that were used to determine wood properties for comparison. No significant differences were determined for wood property data at BH, MID, or 3Q. However, data at 1Q showed higher ring SG (p . 0.043) and percent latewood (p . 0.018) for the northern side, although no differences were observed in the earlywood or latewood SG. This indicated that the higher ring SG for the northern direction results from a greater proportion of latewood. Partitioning the data into estimated juvenile-transition wood and mature wood zones demonstrated that the greater ring SGand percent latewood values in the northern direction occurredwithin the mature wood zone. Findings presented herein appear to provide the first demonstration of variation in wood properties with respect to cardinal direction for a pine species growing in a humid subtropical climate.

Comparison of the Structure of Juvenile and Mature Wood of Larix decidua Mill. from Fast-Growing Plantations in Poland

The main objective of this study was to reveal variations in the wood structure and to define the location of the boundary between juvenile and mature wood Larix decidua Mill. from fast growing tree plantations. To reach these findings, the samples of wood from four short-rotation plantations in Central Europe (Poland) were selected and compared. The procedure used to determine the demarcation point between juvenile and mature wood resulted in strong correlations between the width of growth rings and the initial cambial age. It was found that the structure of wood from fast-growing trees differs from commercially available timber. The wood derived from plantations revealed wider annual rings than wood from natural forests. The analysis of the variability of wood structure resulted in defining the location of the boundary between juvenile and mature wood. Based on the analysis of structure of annual rings, the boundary of juvenile wood in fast-growing larch is placed between the 12th and 15th annual ring. The investigated material contained high proportions of juvenile wood, which makes up more than 70% of trunk volume. On the microscopic level, no visible changes, indicating the presence of a mature wood zone, were found.

Anatomical properties of Eucalyptus grandis wood and transition age between the juvenile and mature woods

The present study is aimed at evaluating the anatomical characteristics of Eucalyptus grandis wood and the transition age between the juvenile and mature woods. Four trees of 23-year-old E. grandis were used. The juvenile, transition and mature wood zones were found by fiber length measurement from the pith to the bark. Samples were taken from juvenile wood, transition wood and mature wood and the anatomical properties of the wood were determined. The results showed that: (1) the fiber length, cell wall thickness, slenderness ratio, cell wall fraction and tangential vessel diameter increase from pith to bark; (2) fibers, axial parenchyma, ray parenchyma cells and vessels are equally distributed from pith to bark; (3) the transition zone between juvenile and mature wood occurs between the age of 8 and 13 years.

Effect of early release from intense competition within high density natural regeneration on the properties of juvenile and mature wood of 40-year-old Sitka spruce (Picea sitchensis (Bong.) Carr.)

While numerous studies have considered the effects of initial planting spacing on wood properties, little is known about the properties of trees grown from dense natural regeneration and the effects of early release from competition. This study examined 40-year-old Sitka spruce (Picea sitchensis (Bong.) Carr.) trees originating from regeneration with an initial density ∼27,000 stems ha−1. Treatments were a control of natural regeneration with no interventions and regeneration re-spaced to two metres (height ∼1.6 m). Density and strength measurements were made on the juvenile and mature wood. No differences were observed in density, modulus of elasticity (MOE) and modulus of rupture (MOR) between the juvenile and mature wood of the control trees. By contrast, the juvenile wood of re-spaced trees was less dense and had a lower MOE than the mature wood. The control trees revealed a uniformity of properties across the juvenile and mature wood zones, unlike the re-spaced treatment. Re-spaced trees appear to adapt to their new environment by producing more thin-walled conductive tissue and lowering overall cell wall density that continues from the juvenile to mature wood. Higher compression wood levels in the control trees are thought to be linked to stem slenderness.

Determination of fiber length and juvenile and mature wood zones in Acer velutinum Boiss. trees grown in Iran

In our investigation we studied fiber lengths and the transition age from juvenile to mature wood in Acer velutinum Boiss. For this purpose, samples from three normal maple trees at a Noshahr site in northern Iran were selected. Disks were cut at breast height. Test samples were taken along a radial direction from the pith to the bark, accounting for every ring during a 48-year period. We used the Franklin method to distinguish between fibers of juvenile and mature wood. The results show that the fiber length increased along the radial direction from the pith to the bark. The transition age between juvenile and mature wood was determined at the 14th annual ring from the pith.

Determination of fiber length and juvenile and mature wood zones from Hevea brasiliensis trees grown in Brazil

The aim of this study was to verify the average fiber length and the juvenile and mature wood zones from Hevea brasiliensis proveniences from a reforestation area in Sao Paulo, Brazil. For this purpose, five Hevea brasiliensis trees were randomly collected from a 50-year-old plantation, located in Balsamo, Sao Paulo, Brazil. The trees were cut and five flat sawn boards were obtained. The juvenile and mature wood zones were determined by fiber length measurement from the pith to the bark. The results showed that: (a) the juvenile wood of this species occurred approx. between 40 and 55 mm, from the pith, and from this point forward, the mature wood zone was found; (b) there was a significant difference between the average fiber length of juvenile wood (1.26 mm) and mature wood (1.51 mm).

The relationship between the form of dead bark and lignin content in Scots pine (Pinus sylvestris L.)

In this study the proportion of lignin in wood tissue was analysed in 36 Scots pines differing in their form of dead bark. Material for analyses came from pure pine stands aged 89-91 years located in northern Poland. Chemical analyses were performed in the mature wood zone comprising the last 10 diameter increments located at a height of 1.3 m (DBH). Results indicate differences in lignin content and growth rate in pines whose stems were characterized by different forms of dead bark. The study showed significant differences in lignin contents between trees with shell and scaly dead bark and trees with ropy bark form. Pines with ropy bark were characterized by the lowest lignin content in relation to trees having the other 2 forms of dead bark. Moreover, a relationship was observed between diameter growth rate and lignin content in wood tissue.

Mold resistance of medium density fiberboard panels made from black spruce, hybrid poplar, larch and a mixture of S-P-F chips

Eleven types of medium density fiberboard (MDF) panels were manufactured from fresh-felled trees of black spruce, hybrid poplar, larch and a mixture of spruce, pine and fir (S-P-F) wood chips at a target density of 740 kg/m3 bonded with 10% commercial urea-formaldehyde (UF) resin. Black spruce panels were fabricated from fibers located in three wood age zones (1–20, 21–40 and over 40 annual rings) of bottom logs, as well as top, middle and bottom logs, separately. Hybrid poplar panels were made from three clones with codes 915303, 915311, and 915313. Mold resistance of these panels was tested by exposing panels in an incubation chamber set at favourable temperature and relative humidity (RH) conditions for mold growth for 4 weeks. The results indicate that MDF panels made from black spruce juvenile wood (1–20 annual rings) were more resistant to mold infection comparing to the other types of panels based on Duncan’s multiple-range test at 0.05 of probability. Panels made of fibers taken from black spruce transition zone (21–40 annual rings) and mature wood zone (over 40 annual rings), and top, middle and bottom logs, hybrid poplar clones 915303, 915311, and 915313, larch and S-P-F wood chips were not significantly different, and these panels were severely infected by mold after 4 weeks of incubation. Nine fungal species predominately infected the tested samples. This laboratory experiment indicates that MDF panels made from black spruce transition and mature wood, and top, middle and bottom logs, hybrid poplar, larch and a mixture of S-P-F chips are susceptible to mold growth and protection measures are necessary in order to improve mold susceptibility of those panels when used in humid conditions or for exterior applications.

Propriedades de resistência e rigidez da madeira juvenil e adulta de Pinus taeda L.

O objetivo do presente trabalho foi o estudo da variabilidade das propriedades de resistência e rigidez à flexão estática e à densidade aparente (12%) entre a madeira juvenil e adulta de Pinus taeda L., de 37 anos de idade, procedente do Horto Florestal de Manduri, Estado de São Paulo. Na primeira parte do trabalho foram determinadas a região de madeira juvenil, a região de transição e a região de madeira adulta, por meio de estudos anatômicos (comprimento dos traqueídes axiais), segundo as recomendações das normas ABNT e IAWA. Os resultados mostraram que a região de madeira juvenil dessa espécie ocorre aproximadamente até o 18º anel de crescimento. Na segunda parte do trabalho foram analisados a resistência (módulo de ruptura - MOR) à flexão, o módulo de elasticidade (MOE) nessa mesma solicitação e a densidade aparente (12%) para as madeiras juvenil e adulta. Os resultados mostraram que o MOE e o MOR da madeira juvenil foram menores e mais variáveis que aqueles obtidos para madeira adulta. A densidade apresentou a mesma tendência observada nas propriedades avaliadas no ensaio de flexão estática.

Basic wood properties of European larch from fast-growth plantations in eastern Canada

A number of basic wood quality characteristics were evaluated in 10 European larch (Larixdecidua Mill.) trees, 5 from each of two rapidly growing plantations in eastern Ontario (approximately 30 years old) and western Quebec (approximately 25 years old). Characteristics evaluated included growth rate, relative density, fibre dimensions, longitudinal shrinkage, alcohol–benzene and water-soluble extractives, and Klason lignin. Radial and longitudinal patterns of variation in wood characteristics were examined and particular attention was given to the distribution of juvenile wood within the stems. Extractive contents were more closely related to the extent of heartwood and sapwood in the stems than to differences between juvenile and mature wood. Relative density was generally lower at the centre of the stem than in the mature outer wood, but the change was usually gradual, with no distinct boundary between juvenile and mature wood zones. The most useful characteristic for delimiting the juvenile wood zone appeared to be longitudinal shrinkage. On this basis, the transition from juvenile to mature wood occurs at about 15 rings out from the pith at breast height. Ranking of individual trees for wood density was judged to be reliable at about 7 years of age at this height level.

Variation in tracheid length in blue pine (Pinus wallichiana A. B. Jackson)

Radial pattern of variation in the first-formed earlywood tracheid length in blue pine shows that it increases outwards from the pith at first rapidly up to the 10th ring, then more slowly up to about the 40th ring and thereafter it remains more or less constant. It is correlated significantly and positively with both age and distance from the pith up to 40 years. In the mature wood zone i.e. after 40 years, tracheid length is not affected by either age or distance from the pith. The mean tracheid length of mature wood is significantly and positively correlated with the mean tracheid length of juvenile wood and also with the tracheid length of each of the juvenile wood rings. Based on the results of this study some guidelines for sampling are proposed. Trees with greater than 51 cm d.b.h. or above 40 years of age can be compared by sampling only four outermost consecutive mature wood annual rings; and those with less than 51 cm d.b.h. or below 40 years of age can be compared by sampling any of the juvenile wood rings.