Levels Of Coarse Woody Debris Research Articles

Prescribed fire is an effective restoration measure for increasing boreal fungal diversity.

Intensive forestry practices have had a negative impact on boreal forest biodiversity; as a consequence, the need for restoration is pressing. Polypores (wood-inhabiting fungi) are key decomposers of dead wood, but, due to a lack of coarse woody debris (CWD) in forest ecosystems, many species are under threat. Here, we study the long-term effects on polypore diversity of two restoration treatments: creating CWD by felling whole trees and prescribed burning. This large-scale experiment is located in spruce-dominated boreal forests in southern Finland. The experiment has a factorial design (n = 3) including three levels of created CWD (5, 30, and 60 m3 ha-1 ) crossed with burning or no burning. In 2018, 16 years after launching the experiment, we inventoried polypores on 10 experimentally cut logs and 10 naturally fallen logs per stand. We found that overall polypore community composition differed between burned and unburned stands. However, only red-listed species abundances and richness were positively affected by prescribed burning. We found no effects of CWD levels created mechanically by felling of trees. We show, for the first time, that prescribed burning is an effective measure for restoring polypore diversity in a late-successional Norway spruce forest. Burning creates CWD with certain characteristics that differ from what is created by CWD restoration by felling trees. Prescribed burning promotes primarily red-listed species, demonstrating its effectiveness as a restoration measure to promote diversity of threatened polypore species in boreal forests. However, because the CWD that the burning creates will decrease over time, to be functional, prescribed burns need to be applied regularly on the landscape scale. Large-scale and long-term experimental studies, such as this one, are invaluable for establishing evidence-based restoration strategies.

Reduced forest vulnerability due to management on the Hualapai Nation

Tribal nations in the US have worked to uphold their long history of managing forests in ways that reduce fuels, support ecosystem functioning, and enhance Indigenous livelihoods. Forests on the Hualapai Nation at the western end of the Grand Canyon have been actively managed for decades using fire and other treatments. We collected data on tree size and age structure, forest understory characteristics, and surface fuels to explore how the legacy of forest management, historical surface fire, and recent prescribed fire, have influenced the contemporary structure of the ponderosa pine-Gambel oak forest on the Hualapai tribal lands (hereafter Hualapai forest). Current overstory tree density (range: 361.1 to 1664.0 trees ha−1) and basal area (16.0 to 29.8 m2 ha−1) place the Hualapai forest in an intermediate state among more open vs. denser southwestern forests, but additional characteristics set the Hualapai forest apart from others in the region. In particular, diameter distributions of live ponderosa pine in the Hualapai forest are dominated by mid-diameter trees (trees ∼18-33 cm dbh), while diameter distributions of live ponderosa pine on forest lands across Arizona indicate a higher number of small-diameter trees (trees <23 cm dbh) and a lower number of large-diameter trees. This finding of a relatively lower number of small-diameter trees, which serve as ladder fuels, in the Hualapai forest may indicate better protection against severe wildfire, although some individual sites might be at greater risk due to higher numbers of small live trees and snags, seedlings and saplings, along with higher levels of coarse woody debris. Beyond fire risk, the Hualapai forest is lacking in large live trees and snags that support cultural needs and sustain its ecological functioning. Current management approaches are helping support the recovery of large trees and snags. While the Hualapai forest is at risk due to its location at the lower elevational limit of ponderosa pine in the Southwest, the legacy of past management and prescribed fire has differentiated it from similar forests in the region. Continued management informed by Indigenous perspectives can best position the Hualapai forest to sustain its structure and function as climate warms as well as achieve socio-cultural and ecological outcomes most important to the people of the Hualapai Tribe.

Response of terrestrial salamanders to the decade following timber harvest in hardwood forests

Timber harvesting can have strong effects on terrestrial salamanders, which are critical components of forest ecosystems and indicators of environmental change. Effects of harvest methods such as clearcutting have been studied in the short term, but few studies have examined salamander trends throughout the decade following timber harvest. The effects of other harvest methods, such as patch cuts and shelterwoods, also remain unclear. We examined the effects of clearcuts, patch cuts, and shelterwood harvests on salamander relative abundance one year before and up to eleven years after harvest in clearcuts, shelterwoods, and patch cuts at the Hardwood Ecosystem Experiment (HEE). A total of 41,858 salamanders representing ten species were captured under artificial coverboards over this period with eastern red-back (Plethodon cinereus) and zigzag (Plethodon dorsalis) salamanders dominating. No significant declines occurred in the first three years following harvest, but salamander captures declined in the 4–6 and 7–11 year periods in clearcuts and patch cuts, and no signs of post-harvest recovery were found. Rapid vegetation recovery and high volumes of coarse woody debris (CWD) following harvest likely protected the microclimate used by salamanders; when CWD levels dropped 4–6 years after harvest, a corresponding decline in salamander captures occurred. These years also corresponded to a regional drought that likely increased temperature and decreased moisture in harvest openings, further impacting terrestrial salamanders. Neither the first nor second shelterwood harvests produced declines in salamanders, suggesting that canopy retention could prevent declines due to changing microclimate or drought. Our results highlight the need to consider compound effects of disturbances such as drought and canopy loss and the importance of long-term salamander monitoring following timber harvest to capture lagged effects and eventual recovery.

The effects of harvest frequency on coarse woody debris and its use by fauna

Abstract Context Forest harvest practices have the potential to alter habitat available to forest fauna. One aspect of harvesting practice that has received little research attention is the effect of multiple rotations on available habitat, and its use by fauna. Aims We examined one component of habitat that is critical to many taxa, coarse woody debris (CWD). In forests of north-eastern New South Wales, Australia, we aimed to assess whether harvest (moderate intensity for saw-logs) frequency influences CWD levels, and whether fauna activity on CWD varies with harvest frequency and with CWD attributes. Methods Using habitat surveys and camera trapping, we compared CWD quantity and quality in unharvested sites with those that had been harvested once, twice and three times, and assessed fauna use of CWD. Key results We found significantly greater volume of non-hollow CWD, and of medium-sized pieces (15–30 cm diameter), in sites harvested three times versus unharvested sites. Camera trapping showed that the activity and composition of fauna were similar across these harvest treatments. Our results also found that a broad range of fauna used a variety of logs, including large and small logs in various states of decay. Conclusions Consistent with other Australian studies, our results suggested that there are no detectable impacts of harvesting on hollow or large CWD. Further studies using radio-tracking or similar methods are needed to more comprehensively assess how CWD is used by fauna. Implications CWD was used broadly by a range of species and should be targeted for retention under ecologically sustainable forest management practices to ensure that it continues to be present in harvested landscapes. This is particularly important in places where an intensification of practices is being considered.

Effects of species and environmental factors on browsing frequency of young trees in mountain forests affected by natural disturbances

We analyzed the effects of natural disturbances on relationships between forest regeneration and ungulate browsing. Specifically, we asked if natural disturbances affect preferences of ungulate herbivores towards tree species and size classes; and how local and landscape-scale environmental factors affect spatial patterns of ungulate browsing in disturbed forest ecosystems. Our research was conducted in the Tatra National Park, a mountain range affected by large-scale intense disturbances over the last few decades. We used field data collected in a network of regularly distributed 600 circular sample plots where young trees were recorded and examined for browsing signs. Using forage ratio, the Ivlev’s electivity index and chi square tests, we found that the preferences of wild ungulates were similar to those known from undisturbed forests. Our analyses showed preference for deciduous species, especially rowan, sycamore maple and goat willow. The most common species, Norway spruce was strongly avoided by browsing ungulates. Short saplings from 0.5 m to 1.3 m tall were the most browsed size class. Effects of environmental variables on browsing were analyzed with zero-inflated negative binomial models. The occurrence of browsing was positively related to the total number of young trees and mature tree mortality. This suggests the influence of disturbances on spatial patterns of browsing pressure. The browsing frequency (number of browsed individuals per plot) was affected by mature tree mortality and densities of young palatable trees, including silver fir, sycamore maple and rowan. Elevation, geology, soil type, and potential solar radiation did not show any significant influence on the browsing frequency. Browsing frequency was also positively related to the distance to roads and hiking trails, indicating the role of human pressure upon the behaviour of ungulate herbivores. In addition, we analyzed the effects of accumulation of coarse woody debris (CWD) on browsing frequency and found this relationship to be nonlinear. Analysis for all tree species revealed that at low levels of CWD, the browsing frequency increased with CWD accumulation as a consequence of forest regeneration. However, a large amount of CWD (>25 m3) resulted in a decrease in browsing frequency, and this relationship was species-specific.

Benchmarks and predictors of coarse woody debris in native forests of eastern Australia

AbstractFallen coarse woody debris (CWD) is critical to forest biodiversity and function. Few studies model factors that influence CWD availability, although such investigations are critically needed to inform sustainable forest management. We assess benchmark levels of CWD in unharvested native forests and those harvested for timber, across a range of forests in north‐east New South Wales, Australia. We found timber‐harvesting was the dominant driver of CWD, with almost double the count (pieces ha−1) and volume (m3 ha−1) of total CWD in selectively harvested than unharvested sites. This pattern was consistent across wet and dry forest types. Harvested sites had greater counts of hollow‐bearing logs, and greater volumes of small and medium‐sized CWD (15–50 cm diameter) than unharvested sites. There was no effect of harvesting on the volume of large CWD (&gt;51 cm diameter). Total volumes of CWD (&gt;15 cm diameter) varied from 114 to 166 m3 ha−1. We found few differences in CWD counts and volumes between forest types, with grassy woodlands and forests containing less CWD than other dry and shrubby forest types, reflecting lower potential input rates. The CWD levels recorded here are similar to those recorded in dry and wet sclerophyll forests elsewhere in Australia and are typical of global estimates for ‘old growth’ forests. Using general linear models we captured up to 57% of the variation in CWD across sites, and found that timber harvesting, topography and the numbers of standing hollow‐bearing and dead trees were significant predictors of CWD. Values for unharvested forest provide a benchmark that could be used to inform retention guidelines for CWD in managed forests in this region. Further assessment of the effect of repeat timber harvesting is needed to fully understand its impact on CWD dynamics, especially if forest residues resulting from timber harvesting are removed from native forests for bioenergy production.

Net CO2 emissions from a primary boreo-nemoral forest over a 10 year period

Primary forests play an important role in the global carbon balance. With little to no human intervention, primary forests are shaped and characterised by disturbances such as weather extremes, fire, insect and pathogen attacks. Such disturbances have a direct impact on the volume of coarse woody debris (CWD) which contributes to the total ecosystem respiration (Re). There are currently few studies that present continuous long term measurements of the carbon balance of northern primary forests. We used the eddy covariance method to measure continuous carbon dioxide (CO2) fluxes from a Swedish primary boreo-nemoral forest over a ten year period. By mapping the measured CO2 fluxes to the forest ecosystem we could indicate that small areas that had some form of disturbance and areas with significant levels of CWD within the eddy covariance footprint contributed to the total Re resulting in the forest being a net carbon source. A weighing algorithm was used to account for directional ecosystem heterogeneity and to estimate a representative carbon balance for the ecosystem. The forest ecosystem was a continuous source of CO2 to the atmosphere, losing around 25Mg per hectare of CO2 over a ten year period.

Long‐Term Soil Changes from Forest Harvesting and Residue Management in the Northern Rocky Mountains

Core Ideas Long‐term (38 yr) soil changes after woody residue management were evaluated. There were no differences in soil physical and chemical properties. Differences in extractable cation pools may be due to different vegetation litter inputs. Soil changes associated with forest harvesting, differing utilization levels, and post‐harvest prescribed burning were determined using an empirical study to investigate the long‐term impacts on soil physical and chemical properties at Coram Experimental Forest in northwestern Montana. In 1974, two replications of three regeneration cuttings (shelterwood, group selection, and clearcut) were installed. In addition, four residue management regimes (high utilization with no burning, medium with no burning, medium with broadcast burning, and low with broadcast burning) were implemented (∼74, 63, 65, and 54% wood removal, respectively). Thirty‐eight years after harvesting, changes were evaluated in mineral soil and forest floor physical and chemical properties (organic matter [OM], C, N, Ca, K, and Mg pools, soil bulk density, and pH) and in coarse woody debris levels. There were no differences in soil pH and bulk density across all regeneration cuttings and residue treatments, probably due to the minimal soil effects associated with the forest harvesting operations that were used (hand felling and cable yarding). Comparisons between harvest and burning and the control indicate no statistical differences in OM, C, and N contents. Minor differences in extractable cation pools were noted in several comparisons among the treatments; these may be attributed to litter inputs from the differing vegetation compositions of overstory and shrub layers rather than nutrient changes within the mineral soil itself. At this moist‐cool forest, intensive biomass utilization, with or without broadcast burning, had few long‐term impacts on soil properties of soil C, OM, and nutrients.

Impact of wind-induced microsites and disturbance severity on tree regeneration patterns: Results from the first post-storm decade

In two hemiboreal mixed spruce–hardwood forests in north-east Estonia, we studied (1) which factors affect tree regeneration survival and development during the first post-storm decade and (2) how these effects change in time. Regeneration height and mortality of the tree species black alder (Alnus glutinosa (L.) J. Gaertn.), birch (Betula pendula Roth., Betula pubescens Ehrh.), Norway spruce (Picea abies (L.) Karst.) and European rowan (Sorbus aucuparia L.) were analysed in moderately and heavily damaged stands, in two types of windstorm-created microsites, i.e. root-plate pits and mounds of uprooted trees, and on intact soil at different stages since disturbance.Regeneration was significantly taller in heavily damaged areas and species traits regarding tree height only became noteworthy at later stages since disturbance. Mortality probability was initially indifferent to microsite type and increased later for regeneration on intact soil compared to regeneration on the storm-induced microsites. Mortality increased with storm severity for A. glutinosa and Betula, whereas mortality of P. abies was initially low and became higher with time since disturbance in areas with increased levels of coarse woody debris. Eventually, height and height increment in previous years were clearly negatively related to mortality probability and competition levels in previous years increased chance of death. The relatively high spatial heterogeneity and trends in dominance of post-storm microsites by different tree species increase disturbance-emulating management options. In conclusion, regeneration mortality and species composition are initially directed by exogenous factors linked to storm severity and microsite heterogeneity, generating a degree of spatial partitioning within a microsite, whereas gradually species’ life-history traits and competition take over.

The effects of different types of management, functions, and characteristics of stands in Polish forests on the amount of coarse woody debris

In the modern forestry paradigm, many factors influence the amount of coarse woody debris (CWD). The present paper analyzes the effects of both local (national) programs (special functions of forests) and European programs (Natura 2000 sites), as well as the individual characteristics of forest stands. The study was conducted on 2,752 sampling plots distributed over an area of about 17,500 ha and located in lowland stands having a species composition typical of large areas in central Europe. Natura 2000 areas contained significantly more CWD (8.4 m3/ha) than areas not covered by the program (4.8 m3/ha). However, this is due to the fact that Natura 2000 sites involve well-preserved forest areas, such as nature reserves (26.6 m3/ha). In the managed forests that have been covered by the Natura 2000 program over the past several years, the volume of CWD has not increased. Forests with ecological and social functions differed slightly in the amount of CWD. More CWD occurred in protected animal areas (8.7 m3/ha) than in stands damaged by industry (3.9 m3/ha). Intermediate CWD levels were found in water-protection forests and in forests located around cities and military facilities. In managed forests, the lowest CWD volume was observed in middle-aged stands. The species composition of the stand had little effect on the volume of CWD. Only stands with a predominance of ash and alder had higher CWD levels (13.5 m3/ha). More CWD was found in stands whose species composition did not represent the potential site quality (6.4 m3/ha) than in habitats with the optimum species composition (3.8 m3/ha). CWD volume should be systematically increased taking into consideration local natural conditions. Such efforts should be focused on particularly valuable regions, and especially on Natura 2000 sites, where the threshold values reported from other European forests should be reached. Leaving some trees to die naturally and retaining reasonable amounts of such trees ought to be incorporated into CWD management practice in Poland.

Does coarse woody debris density and volume influence the terrestrial vertebrate community in restored bauxite mines?

Coarse woody debris (CWD) is a critical functional and structural component of forest and woodland ecosystems, providing habitat for many species, and is an important consideration in forest and woodland restoration. CWD is very slow to develop naturally so, to accelerate the return of CWD-dependent species to restored areas, CWD is commonly returned manually. However, few studies have tested the effectiveness of such a strategy. We investigated whether the provision of CWD, heaped into ‘habitat piles’ of varying density (0.4–5.7pilesha−1), was effective in accelerating recolonisation by reptiles, frogs and mammals into 3-year old restored bauxite mine-pits in south-western Western Australia. Both reptile and mammal communities, and the abundances of some individual species, differed significantly between unmined and restored forest but the provision of CWD had only a weak effect in accelerating recolonisation. Acritoscincus trilineatus abundance showed a weak positive relationship with habitat pile density and both Cryptoblepharus buchananii and Christinus marmoratus, species that are very rare in restoration, were recorded adjacent to habitat piles in two and one mine-pits respectively. The weak effects of CWD in accelerating recolonisation were likely due to the differences in vegetation between unmined and restored forest, resulting in restored forests being primarily inhabited by generalist species that did not require CWD, and the highest habitat pile densities being ⩽6% of log densities in unmined forest, suggesting that CWD-dependent species perceived all mine-pits as having similarly low levels of CWD, compared to unmined forest. Our results suggest that the provision of CWD in restored areas is critical to accelerate recolonisation of CWD-dependent fauna, but this will require consideration of both CWD spatial connectivity and temporal continuity. Spatial connectivity would be best achieved through CWD densities that approximate those in reference ecosystems, whereas temporal continuity will be harder to achieve, particularly in systems where CWD is slow to develop, and will require the development of innovative techniques and long-term management. However, ensuring the spatial connectivity and temporal continuity of CWD in restored areas should greatly increase their biodiversity value.

Pre-wildfire fuel reduction treatments result in more resilient forest structure a decade after wildfire

Increasing size and severity of wildfires have led to an interest in the effectiveness of forest fuels treatments on reducing fire severity and post-wildfire fuels. Our objective was to contrast stand structure and surface fuel loadings on treated and untreated sites within the 2002 Rodeo–Chediski Fire area. Data from 140 plots on seven paired treated–untreated sites indicated that pre-wildfire treatments reduced fire severity compared with untreated sites. In 2011, coarse woody debris loading (woody material&gt;7.62cm in diameter) was 257% higher and fine woody debris (woody material&lt;7.62cm) was 152% higher on untreated sites than on treated sites. Yet, in spite of higher levels of coarse woody debris on untreated sites, loadings did not exceed recommended ranges based on published literature and many treated sites fell below recommendations. By 2011, basal area and stand density on treated sites and stand density on untreated sites met management guidelines for ponderosa pine forests, but untreated sites had basal areas well below recommendations. Snags declined over this period and only three plots had snags that met minimum size and density requirements for wildlife habitat by 2011. The effects of pre-wildfire treatments are long-lasting and contribute to changes in both overstorey and understorey fuel complexes.

A simplified inventory approach for estimating carbon in coarse woody debris in high-biomass forests

Forests carrying large quantities of live and dead wood are important carbon (C) stores. Here, we investigate how the inventory of coarse woody debris (CWD) and its embedded C (CWD-C) may be designed efficiently at the scale of logs, plots, and the landscape in Tasmanian tall Eucalyptus obliqua forests, which have very high levels of CWD (here 375–1085 m³ ha–1). From a set of 12 sites representing different times since disturbance, a thorough census of dead wood >10 cm in diameter was carried out at five sites using a fixed-plot (50 × 50 m) approach. This showed that 90% of the volume can be captured by recording only CWD logs >40 cm in diameter. Based on this approach and on the known density and C content of five different decay-classes, volume, mass, and CWD-C was determined for all 12 sites. To obtain an accurate estimate of CWD-C at the landscape scale, it was found to be sufficient to allocate entire individual logs to single decay-classes and to use one global value for C content instead of decay-class-specific values. The most decayed logs, which are difficult to measure, could be ignored. However, at the plot level, no relationships were found between CWD mass and either standing or downed CWD or standing-tree biomass, limiting the utility of these proxies for assessing CWD volume.

Regeneration responses to gap size and coarse woody debris within natural disturbance-based silvicultural systems in northeastern Minnesota, USA

The use of silvicultural systems based on the natural disturbance patterns for a given region has been suggested as an approach for promoting late-successional forest characteristics and maintaining native biodiversity in managed forests. To this end, we examined the effectiveness of natural disturbance-based harvest gaps in maintaining and restoring native tree species diversity within second-growth northern hardwood systems in northeastern Minnesota, USA. In particular, tree regeneration and downed coarse woody debris (CWD) were measured 6- and 7-years post-treatment in 46 gaps that emulated the historic range of natural canopy opening sizes for northern hardwood forests in the upper Great Lakes region. Measurements were compared across gap size-classes (>0–0.02, >0.02–0.04, >0.04 ha) and with closed canopy portions of these systems. In addition, the factors affecting the recruitment of Betula alleghaniensis across gaps and within closed canopy were investigated due to the historical importance of this species in these forests. Seedling and sapling densities increased in harvest gaps; however, results indicated that these gaps did little to increase tree diversity, including the recruitment of shade mid-tolerant species, such as B. alleghaniensis. Gaps were dominated primarily by Acer saccharum and age distributions of dominant saplings within gaps indicated that most individuals established 10–40 years prior to gap formation. Establishment of B. alleghaniensis was strongly related to highly decayed, large coniferous pieces of CWD with little recruitment occurring on the undisturbed forest floor. Age distributions of B. alleghaniensis established on CWD suggest this species can persist on this substrate for at least a decade. Levels of CWD increased with increasing gap size; however, all gaps had lower levels of CWD compared to the surrounding intact forest. Our results suggest that management regimes based on natural canopy gap sizes within northern hardwood systems will do little to restore native tree diversity if provisions for creating the suitable seedling microsites historically generated by natural disturbance (e.g., exposed mineral soil, highly decayed wood) are not included within management prescriptions.

Biofuel harvests, coarse woody debris, and biodiversity – A meta-analysis

Forest harvest operations often produce large amounts of harvest residue which typically becomes fine (foliage, small limbs and trees) and coarse woody debris (snags and downed logs). If removed at harvest, residual biomass has potential to be a local energy source and to produce marketable biofuel feedstock. But, CWD in particular serves critical life-history functions (e.g., breeding, foraging, basking) for a variety of organisms. Unfortunately, little is known about how forest biodiversity would respond to large scale removal of harvest residues. We calculated 745 biodiversity effect sizes from 26 studies involving manipulations of CWD (i.e., removed or added downed woody debris and/or snags). Diversity and abundance of both cavity- and open-nesting birds were substantially and consistently lower in treatments with lower amounts of downed CWD and/or standing snags, as was biomass of invertebrates. However, cumulative effect sizes for other taxa were not as large, were based on fewer studies, and varied among manipulation types. Little is currently known about biodiversity response to harvest of fine woody debris. Predicting the effects of biomass harvests on forest biodiversity is uncertain at best until more is known about how operational harvests actually change fine and coarse woody debris levels over long time periods. Pilot biomass harvests report post-harvest changes in CWD levels much smaller than the experimental changes involved in the studies we analyzed. Thus, operational biomass harvests may not change CWD levels enough to appreciably influence forest biodiversity, especially when following biomass harvest guidelines that require leaving a portion of harvest residues. Multi-scale studies can help reduce this uncertainty by investigating how biodiversity responses scale from the small scale of manipulative experiments (i.e., 10-ha plots) to operational forest management and how biodiversity response to CWD levels might vary at different spatial and temporal scales and in different landscape contexts.

Decomposition and Fragmentation of Coarse Woody Debris: Re-visiting a Boreal Black Spruce Chronosequence

We re-visited a seven-stand boreal chronosequence west of Thompson, Manitoba, Canada, in which coarse woody debris (CWD) and its instantaneous decomposition were measured in 2000. New CWD measurements were performed in 2007, and tree inventories updated to provide mortality and snag failure data. These data were used to model CWD changes, compare methods of estimating decomposition, and infer possible fragmentation rates. Measured CWD was between 9.7 (in both the 77- and 43-year-old stands) and 80.4 (in the 18-year-old stand) Mg ha−1 in 2007. Spatial variability was high; at most stands CWD levels had not changed significantly from 2000 to 2007. Tree mortality was a significant flux only in older stands, whereas snag fall rate varied by an order of magnitude, from 2.9% y−1 (0.2 Mg ha−1 y−1) in the 9-year-old stand to 9.8% y−1 (2.3 Mg ha−1 y−1) in the 12-year-old stand. A one-pool model based on these inputs underestimated actual 2000–2007 CWD decomposition in the younger stands, suggesting that fragmentation could be an important part of the carbon flux exiting the CWD pool. We compared three independent measures of annual decomposition (k): direct measurements of CWD respiration, rates based on the 7-year re-sampling effort described here, and rates inferred from the chronosequence design itself. Mean k values arrived at via these techniques were 0.06 ± 0.03, 0.05 ± 0.04, and 0.05 ± 0.05 y−1, respectively. The four-pool model suggested that the transition rate between decay classes was 0.14–0.19 y−1; the model was most sensitive to initial CWD values. Although the computed k values implied a problem with chronosequence site selection for at least one site, the overall CWD trend was consistent with a larger number of sites surveyed in the region.

Using field survey and remote sensing to assess rainforest canopy damage following Cyclone Larry

Abstract We surveyed canopy damage in upland and lowland rainforests following Cyclone Larry, which severely impacted the Innisfail and Atherton Tableland regions in March 2006. An existing damage‐classification was used as a basis for field assessment of rainforest canopy damage. Our field measurements showed that the damage categories were not clearly separated. Upland and lowland sites significantly differed in tree fall variables and measures of forest structure. There was a difference in recruitment of disturbance indicator species owing to varying levels of coarse woody debris at sites. Aspect was not found to be a significant variable in predicting damage owing to complexity of topography and the cyclone wind field. Analysis of remotely sensed imagery indicated that only high damage levels could be reliably discerned. Areas of very rapid vegetation growth in severely damaged sites are most easily detected with vegetation indices based on both near infrared and short wave infrared data. Numbers of fallen trees and their trunk orientations can be reliably quantified using high resolution (sub‐metre) colour aerial photography. This permits some estimation of whether the wind field was unidirectional or locally vortical.

Immediate tree uprooting after retention-felling in a coniferous boreal forest in Fennoscandia

We studied the immediate effects of retention-felling on the occurrence of tree uprooting in two different types of boreal spruce forest in Finland to determine whether susceptibility to uprooting is dependent on the biotope. During the first post-treatment year, 7.1% of the trees in the paludified forest type and 1.8% in the upland forest type were uprooted. During the 2 following post-treatment years, uprooting percentages increased considerably (39.3% in the paludified type and 11.5% in the upland type in year 2; 48.3% in the paludified type and 15.2% in the upland type in year 3). Norway spruce (Picea abies (L.) Karst.), the dominant species in both forest types, was the species most susceptible to uprooting. The high uprooting rate in the paludified groups was probably caused by an interaction between the rocky ground and moist overlying peat layer with the shallow root system of P. abies. As paludified forest biotopes are generally recognized to have high biodiversity, the use of green-tree retention in these biotypes may enhance the continuum and survival of sensitive species. Moreover, because of the high level of uprooting, green-tree retention in paludified forest types can quickly and more naturally help restore levels of coarse woody debris.

Effects of structural complexity enhancement on eastern red-backed salamander ( Plethodon cinereus) populations in northern hardwood forests

Managing for stand structural complexity in northern hardwood forests has been proposed as a method for promoting microhabitat characteristics important to eastern red-backed salamanders ( Plethodon cinereus). We evaluated the effects of alternate, structure-based silvicultural systems on red-backed salamander populations at two research sites in northwestern Vermont. Treatments included two uneven-aged approaches (single-tree selection and group-selection) and one unconventional approach, termed “structural complexity enhancement” (SCE), that promotes development of late-successional structure, including elevated levels of coarse woody debris (CWD). Treatments were applied to 2 ha units and were replicated two to four times depending on treatment. We surveyed red-backed salamanders with a natural cover search method of transects nested within vegetation plots 1 year after logging. Abundance estimates corrected for detection probability were calculated from survey data with a binomial mixture model. Abundance estimates differed between study areas and were influenced by forest structural characteristics. Model selection was conducted using Akaike Information Criteria, corrected for over-dispersed data and small sample size (QAIC c). We found no difference in abundance as a response to treatment as a whole, suggesting that all of the uneven-aged silvicultural systems evaluated can maintain salamander populations after harvest. However, abundance was tied to specific structural habitat attributes associated with study plots within treatments. The most parsimonious model of habitat covariates included site, relative density of overstory trees, and density of more-decayed and less-decayed downed CWD. Abundance responded positively to the density of downed, well-decayed CWD and negatively to the density of poorly decayed CWD and to overstory relative density. CWD volume was not a strong predictor of salamander abundance. We conclude that structural complexity enhancement and the two uneven-aged approaches maintained important microhabitat characteristics for red-backed salamander populations in the short term. Over the long-term, given decay processes as a determinant of biological availability, forestry practices such as SCE that enhance CWD availability and recruitment may result in associated population responses.

Restoring natural characteristics in managed Norway spruce [Picea abies (L.) Karst.] stands with partial cutting, dead wood creation and fire: immediate treatment effects

This study investigated the immediate effects of restorative treatments on stand structure in mature managed Picea abies-dominated stands in southern Finland. The restorative treatments consisted of partial cuttings with three levels of coarse woody debris (CWD; 5, 30 and 60 m3 ha−1) with a constant volume of standing retention trees (50 m3 ha−1), and a burn/non-burn treatment. In addition, burned and unburned reference stands without cutting treatment were assessed. Each stand (n=24) included an upland and a paludified biotope; the paludified biotopes burned only partially. Detailed descriptions of stand structure were made for living and dead tree components before and after restorative treatments. Changes in tree diameter distribution and increases in standing dead tree volume were greatest in the combined burning and cutting treatments. The resulting standing dead wood volume varied greatly (1–52 m3 ha−1) between treatments, and was highest on burned paludified biotopes with 60 m3 ha−1 of CWD. The results indicate that creation of down wood was needed to achieve burning and that it is possible to regulate the severity of burnings through the amount of CWD, according to ecological restoration objectives. The study demonstrates that living and dead tree structural characteristics typical of natural postdisturbance early-successional forests can be rapidly restored to mature managed spruce stands, even when a significant portion of wood volume is harvested. These types of restorative cuttings provide one potential means of conserving biodiversity in managed forests.