Lakeshore Sites Research Articles

The wooden artifacts from Schöningen’s Spear Horizon and their place in human evolution

Ethnographic records show that wooden tools played a pivotal role in the daily lives of hunter-gatherers including food procurement tools used in hunting (e.g., spears, throwing sticks) and gathering (e.g. digging sticks, bark peelers), as well as, domestic tools (e.g., handles, vessels). However, wood rarely survives in the archeological record, especially in Pleistocene contexts and knowledge of prehistoric hunter-gatherer lifeways is strongly biased by the survivorship of more resilient materials such as lithics and bones. Consequently, very few Paleolithic sites have produced wooden artifacts and among them, the site of Schöningen stands out due to its number and variety of wooden tools. The recovery of complete wooden spears and throwing sticks at this 300,000-y-old site (MIS 9) led to a paradigm shift in the hunter vs. scavenger debate. For the first time and almost 30 y after their discovery, this study introduces the complete wooden assemblage from Schöningen 13 II-4 known as the Spear Horizon. In total, 187 wooden artifacts could be identified from the Spear Horizon demonstrating a broad spectrum of wood-working techniques, including the splitting technique. A minimum of 20 hunting weapons is now recognized and two newly identified artifact types comprise 35 tools made on split woods, which were likely used in domestic activities. Schöningen 13 II-4 represents the largest Pleistocene wooden artifact assemblage worldwide and demonstrates the key role woodworking had in human evolution. Finally, our results considerably change the interpretation of the Pleistocene lakeshore site of Schöningen.

Lakeshore areas of conservation interest: Characteristics of nursery areas of the threatened lake‐dwelling grayling, Thymallus thymallus

AbstractResearch‐based knowledge is essential for effective conservation and restoration of threatened aquatic species and habitats. Here, our aim was to gather this knowledge on the lake‐dwelling grayling (Thymallus thymallus), typically a riverine fish. Such atypical populations are particularly vulnerable to anthropogenic impacts, including fishing pressure, climate change, eutrophication and waterway construction, some of which affect especially the early life stages. However, there is little information available to guide management and conservation of grayling in lakes. Accordingly, we assessed characteristics of the nursery areas in the threatened grayling population of Lake Puruvesi (eastern Finland). In particular, we used beach seines in two consecutive years to sample lakeshore sites (including islands) that were a priori presumed suitable for grayling. We assessed the occurrence of grayling fry (larvae and post‐larvae <40 mm in length) regarding depth, year, the site's exposure (fetch), bottom shear stress, substrate coarseness and shoreline's north–south orientation. Overall, we found grayling fry in low numbers at every fourth site, with the sites’ exposure and dominant substrate coarseness being most relevant variables. In particular, more exposed sites (i.e. with higher fetch values) and fine‐grained substrates dominated by sand or gravel had more grayling fry. Average depth, bottom shear stress or shoreline orientation along the north–south axis did not have a significant effect. Together, the results suggest that the most important nursery areas for lake‐dwelling grayling are lakeshore zones that are barren and exposed. Hence, the sites share characteristics with those used for reproduction by the more common riverine grayling. We hope that these findings will facilitate lake‐dwelling grayling's management and conservation efforts.

Using microartifacts to infer Middle Pleistocene lifeways at Schöningen, Germany

While archeologists usually favor the study of large and diagnostic lithic artifacts, this study illustrates the invaluable contribution of lithic microartifacts for interpreting hominin lifeways. Across a 64 m2 area of the Middle Pleistocene lakeshore site of Schöningen 13 II-3 in Northern Germany, we recovered a total of 57 small and micro flint artifacts, four small debris pieces, three natural fragments and three bone retouchers in close association with the skeleton of an extinct Eurasian straight-tusked elephant (Palaeoloxodon antiquus). This area lacks the type of formal knapped stone tools that would normally constitute the focus of archeological interpretations. By adopting a holistic approach, including morpho-technical analysis, experimental archeology, and use-wear and residue analyses, we demonstrate that these small and microartifacts are resharpening flakes that tell the story of the site. Fifteen resharpening flakes preserve microwear traces of processing wood. Microscopic residues of wood adhered to the former working edges of the tools corroborate this observation. Additionally, hominins used a sharp-edged, natural fragment of flint to process fresh animal tissue, which likely originates from the butchery of the elephant. These results provide unique, 300,000-year-old evidence for the functionally interconnected use of lithic, osseous and wood technologies. Furthermore, we document in-situ transformations of stone tools and the presence of both curational and expedient behaviors, thereby demonstrating the temporal depth of hominin activities at the lakeshore where the elephant died, and in the broader landscape as a whole.

The influence of stream tributaries to lakes and upstream forest harvest on benthic, lake delta communities

Lake deltas (LDs) receive nutrient and energy subsidies from upstream habitats. These inputs can support LD benthic production. However, LDs may also receive impacts from upstream disturbance. Small lakes in coastal British Columbia often have inflow streams from areas with forest harvesting, which is pervasive in this region. Forest harvesting impacts can alter stream production, although downstream impacts on lakes, compared to streams, are less studied and understood. We aimed to test whether benthic LD communities (biofilm and invertebrates) are more productive and (or) diverse compared to other communities along the lakeshore (LS) (non-LDs), and whether there are differences in LD communities associated with upstream harvesting. We compared LD communities with upstream riparian areas that were harvested (HLD) or relatively undisturbed (ULD), and LS sites. We found that while community structure was similar across sites, ULDs had higher biofilm and invertebrate standing stocks (1.5- and 2-times higher, respectively) than HLDs and LS sites. Our results suggest subsidies and benthic production were higher at ULDs. These results advance our understanding of forest harvesting disturbance and stream-lake connectivity.

Multiproxy study of 7500-year-old wooden sickles from the Lakeshore Village of La Marmotta, Italy

The lakeshore site of La Marmotta is one of the most important Early Neolithic sites of Mediterranean Europe. The site is famous for the exceptional preservation of organic materials, including numerous wooden artefacts related to navigation, agriculture, textile production, and basketry. This article presents interdisciplinary research on three of the most complete and well-preserved sickles recovered from the site, yet unpublished. All the components of the tools are analysed: the stone inserts, the wooden haft and the adhesive substances used to fix the stones inside the haft. Our innovative methodology combines use-wear and microtexture analysis of stone tools through confocal microscopy, taxonomical and technological analysis of wood, gas chromatography–mass spectrometry analysis of the adhesive substances, and pollen, non-pollen palynomorphs, and phytolith analysis of the remains incorporated within the adhesive. This multiproxy approach provides a significant insight into the life of these tools, from their production to their use and abandonment, providing evidence of the species of harvested plants and the conditions of the field during the harvesting.

A habitat analysis and influence of scale in lichen communities on granitic rock

Environmental factors in saxicolous lichen communities are scale dependent, yet the influence of scale is not often specifically addressed in ecological studies. This study examined three scales [micro (cm), meso (meters–10s of meters), macro (km)] in granite rock communities and compared species assemblages and disturbance regimes in Lake Superior shoreline and inland habitats. Percent cover of lichens and environmental variables were measured in 1280 20 × 20-cm plots across 16 sites (8 lakeshore and 8 inland). A perMANOVA analysis showed that the composition of lakeshore and inland lichen communities was significantly different (F=17.2, df=1,4, P <0.001). Adjacent lakeshore sites were more similar (F=8.550; df = 1, 26; p=0.007) to each other than they were to sites further away, while inland sites were not (F=0.545; df=1, 26; p=0.467). Variation in disturbance is likely more important in determining inter-site variation in inland areas. Mesoscale environmental variables such as solar radiation, height above Lake Superior, and an aspect/slope index were better predictors of species assemblages than microscale variables. However, individual species differed greatly in their associations with specific habitat variables. A host of microhabitats were discovered, with some lichens specializing on rock overhangs, quartz veins, cracks, subtle variations in rock texture, and mafic intrusions within granite.

Neumark-Nord 2 – A multiphase Middle Palaeolithic open-air site in the Geisel Valley (Central Germany)

The lake basin Neumark-Nord 2 (NN 2) is located in the Geisel Valley in Saxony-Anhalt (Germany). It was scientifically investigated between 2003 and 2008. The sediment sequence, which is about 10 to 11m thick, consists of limnic deposits, mostly silts mixed with clays and sands. Sedimentological as well as palynological, malacological and palaeomagnetic investigations, supported by absolute datings, date the entire sequence into the Eemian and Early Weichselian. Within the deposits several archaeological find horizons were discovered. Neumark-Nord 2/2 (NN 2/2) at the beginning of the Eemian and Neumark-Nord 2/0 (NN 2/0) with an early Weichselian were the most important archaeological horizons. Both find horizons represent former lake shore sites, where numerous crushed animal bones and other faunal debris were found, which can be regarded as remains of hunted game. With regard to the rich lithic material, both find horizons differ remarkably. In NN2/2, a simple artefact spectrum with a certain Levallois component and a dominance of denticulated, notched and laterally retouched specimens. In NN 2/0, on the other hand, bifacial tools such as backed knives and sometimes finely retouched scrapers are dominant, while the Levallois component fades completely into the background. Thus, NN 2 is one of the few open-air sites in archaeological landscape of Central German where several middle Palaeolithic find horizons occur in superposition.

Neolithic Bow Case from Lenk, Schnidejoch

The find of a yet unknown type of birch bark container in the site of Schnidejoch (Bernese Alps, Switzerland) was a surprise, for in all the wetland or lakeshore sites of Europe no parallels to this unique object are known to date. First, when only a small part was discovered, it was supposed to have been part of an arrow quiver. However, with the final appearance of the ca. 170 cm long container, it needed a new interpretation. The yew bow found at the same site, 160.5 cm long, would fit perfectly inside. Two silex arrowheads found in the bottom confirm an association with archery. The find could be identified as the first Neolithic bow case.

Flaking stone activity in the tradition of iron smelting from the 8th to 17th centuries AD in the Matano region, South Sulawesi, Indonesia

Lake Matano includes a complex of sites related to an iron-smelting industry that developed at the northeast tip of South Sulawesi between the 8th and 17th centuries AD. The sites include recently discovered underwater sites which possibly submerged because of an earthquake that had occurred in the region. An interesting finding at each site is the presence of flaked artifacts from chert. This raises the question of what was the role of flaked chert in the local tradition of iron smelting. The present study was conducted by combining ethnographic data from Lake Matano and analysis of chert artifacts excavated from the Rahumpu’u lakeshore site. The results show that chert artifacts were used as strike-a-lights and in particular for the combustion of fuel for iron smelting. Flaking chert was one of the skills possessed by the Matano indigenous iron smelters in utilizing the resources in their environment to produce a high quality of iron.

Do still waters run deep? Formation processes of natural and anthropogenic deposits in the Neolithic wetland site Zug‐Riedmatt (Switzerland)

AbstractMicromorphological analysis is used to improve our understanding of wetland site taphonomy and stratigraphy. Twelve profile columns from the Neolithic lakeshore site of Zug‐Riedmatt are macroscopically described here, microscopic analyses are applied to sediment facies, defining 14 sedimentary units (U): The natural basal sediment (U1) is carbonate‐rich, U2 a transition phase to the anthropogenic units U3 to U12. Uppermost are the natural carbonate‐rich U13 and U14. The creation of a micromorphological database and the application of 27 defined indices allow a better understanding of the natural, anthropogenic and taphonomic processes taking place at the site. Natural sediment input originated from Lake Zug, the nearby river Lorze with its delta, and a local marshy waterbody. Organic rich accumulations and calcitic ash played a major role in pH changes, leading to carbonate and silica dissolution, alongside bone alteration. Signs of degradation processes under aerobic and anaerobic conditions within a single layer points to complex, multiphase processes under changing water levels, a milieu belonging to both land and water, an amphibious environment. Finally, good layer preservation seemingly correlates to low aquatic reworking in a stable depositional context linked to the marshy area.

Bats in the changing boreal forest: response to a megafire by endangered little brown bats (Myotis lucifugus)

ABSTRACTWildfire is a pervasive disturbance in the boreal forest, and fire size and severity is increasing due to climate change. Yet, wildfire impacts on boreal bats are unknown. Burned forest may be attractive to bats because of an increase in dead trees and a reduction in vegetative clutter. Using ultrasonic detectors, I sampled little brown bat (Myotis lucifugus) activity at lakeshore and upland sites within burned and mature forest. Activity was greater in mature than burned forest at both upland and lakeshore sites. Activity was greater at lakeshores, where some large snags survived fire. The relative use of edges created by fire, lakeshores and roads was also investigated. Bat activity was greater at lakeshores than other edge types. Bats made little use of burned upland areas or fire-created edges. This study provides a first approximation of the effect of a ‘megafire’ on bats in the boreal forest and suggests that large, severe wildfires may have a negative impact. Protection of mature forest containing large dead trees, particularly along lakeshores, may help to mitigate the loss of habitat due to wildfires. Further work, however, is needed to assess bat activity in burns of differing sizes, severity and age.

Phthalate esters (PAEs) in atmospheric particles around a large shallow natural lake (Lake Chaohu, China)

The pollution of phthalate esters (PAEs) remains an important issue in the world. Current studies mainly focused on atmospheric PAEs in urban area with strong anthropogenic activities, but there were no studies on PAEs in the ambient air around large natural lake. This paper focused on two sites around Lake Chaohu to investigate the monthly occurrence, composition and source of PAEs in the atmospheric particles around large shallow natural lake. New insights into atmospheric PAEs in large shallow natural lake and the overall fate of PAEs in lake ecosystem were given. The concentrations of the Σ13PAEs in atmospheric particles were at a significantly low level ranging from 2740 to 11,890 pg·m−3 and 2622 to 15,331 pg·m−3 in ZM (the lakeshore site) and HB (the downtown site), respectively. There were no statistically significant differences of PAEs between ZM and HB. The highest atmospheric PAE concentrations in August were likely related to the long-range transport from Guangdong Province. Di(2-ethylhexyl) phthalate (DEHP), diisobutyl phthalate (DIBP) and dibutyl phthalate (DBP) were the main PAE congeners. Temporally, DIBP and DBP had the highest fractions in winter and the lowest fractions in summer. It might be justified by the condensation of DIBP and DBP from gas phase to particulate phase at low temperature. Multimedia comparison of PAE profiles in Lake Choahu revealed that low molecular weight (LMW) congeners were transported mainly through water while high molecular weight (HMW) congeners were transported mainly through atmosphere.

Measurement of Astronomical Seeing Using Long Exposure Solar Images

We extend our modified parameter search method (Sridharan, Dashora, and Venkatakrishnan, Solar Phys. 222, 35, 2004) of estimating the Fried parameter ( $r_{0}$ ) from long exposure images to the images obtained from the H $\alpha $ -telescope installed at Merak, a cold desert on the Himalayas, with a modified criterion – the contrast ratio of the observed and the deconvolved images – to identify the true $r_{0}$ . We validated our new method by applying it to the solar granulation data and found that it is as accurate as our modified parameter search method. The median seeing at H $\alpha $ – estimated from the data spanning over 5 months – is ≈ 2 arc-sec and it corresponds to a median $r_{0}$ of 6 cm. About 30% of the estimated values are above $r_{0}=7~\mbox{cm}$ . The diurnal variation of the seeing is consistent with that expected for a lake-shore site. The significance of our method – extracting the underlying seeing from a single long exposure solar image – lies in the fact that it can be easily adopted to monitor seeing with small H $\alpha $ -telescopes that are coveted and eagerly possessed by many solar observatories for patrolling solar flares and filament eruptions.

Hunting, Husbandry, and Human-Environment Interactions in the Neolithic Lakeshore Sites of Western Switzerland

The lakeshores of western Switzerland are one of Europe's best-known Neolithic settlement areas, thanks to dendrochronological dating and the exceptional preservation of organic materials. Against this outstanding background, this study uses zooarchaeological data to answer a series of questions regarding the Neolithic economy, environment and human-environment interactions at these lakeshore sites. It also discusses, within an interdisciplinary framework, the possible impact climatic fluctuations, cultural influence, topographical conditions, and demographic growth had on economic change. The results show that the faunal economy was mainly based on animal husbandry, with fluctuations in the cattle-pig ratio. Hunting also played an important role in the food system and focused mainly on large game, especially red deer, which contributed significantly to the meat supply. The results from comparing these animal bone remains also show that multiple factors, such as topography, climatic conditions, and cultural influence, played a part in the socio-economic organisation of the Neolithic communities. Exploratory procedures such as correspondence analysis support these interpretations.

Sedimentology and micromorphology of the Lower Palaeolithic lakeshore site Marathousa 1, Megalopolis basin, Greece

The present study describes the sedimentology and formation processes of the archaeological site Marathousa 1, part of a Pleistocene lignite-bearing succession at the Megalopolis basin (Southern Greece). The sedimentary sequence of the site comprises about 4–5 m lacustrine and fluviolacustrine clastic deposits found between Lignite Seam II and III. The lower part of this sequence is characterized by relatively high rate subaqueous sedimentation of bedded sands and silts with local evidence of slumping and liquefaction attributed to a seismic event. This part of the sequence contains fluctuating but generally low organic and carbonate content probably associated with cold and arid conditions of a glacial period. The upper part of the sequence follows a major hiatus attributed to exposure and erosion. A series of erosional bounded depositional units are observed in this sequence suggesting important water-level fluctuations. The subaqueous emplacement of the deposits is attributed to subaerial flood-generated, organic- and carbonate-rich dilute mudflows and hyperconcentrated flows. Organic-rich sedimentation culminates with the formation of the overlying lignite seam. This change is interpreted as a gradual return to warmer and humid conditions of an interglacial period.The fossiliferous and cultural levels at Marathousa 1 are associated with the major erosional and exposed surface found in the upper part of the sequence. This surface was part of an extensive mudflat surrounding the lake shore at that time. The overlying mudflows have locally redistributed and buried the archaeological remains, thus preserving this very important butchering site.

A cholera outbreak caused by drinking contaminated water from a fenced lakeshore water-collection site � Kasese District, western Uganda, June 2015

On 20 June 2015, a cholera outbreak affecting > 30 people was reported in a fishing village in Kasese District, western Uganda. We investigated this outbreak to identify the transmission mode and recommend control measures.

Assessment of Wildlife Habitat Attributes at Restoration Projects on Northern Wisconsin Lakeshores

Abstract Housing development along lakeshores typically results in the loss of native shoreline vegetation, which can negatively impact habitat structure and associated wildlife populations. We evaluated vegetation restoration efforts on 2 lakeshores in Vilas County, WI, and contrasted them with undeveloped reference lakeshores. The primary goal of the restoration activities was to restore native understory vegetation and habitat structure. Initial measurements made at reference lakeshores showed greater visual obstruction density, greater sapling and shrub densities, greater abundance of downed woody material, and higher canopy coverage relative to initial measurements made at developed lakeshore sites. Three years post-restoration we observed significant increases in visual obstruction density and increased shrub and sapling density at restoration sites. While restoration of complex understory habitats is a slow and uncertain process, a nonmetric multi-dimensional scaling ordination of wildlife habitat ...

Studying the preservation of plant macroremains from waterlogged archaeological deposits for an assessment of layer taphonomy

Layer taphonomy is one of the major questions in the archaeological research of lakeshore settlements. How fast did these deposits develop? Were they exposed to periodic droughts and decay? Which amount of the originally deposited remains survived until present? Plant macroremains have a great potential as indicators of preservation quality, since they are short-lived and particularly sensible to changes in preservation conditions. This paper reviews previous attempts to use similar proxies to understand layer taphonomy and provides a compilation of almost 50 variables (that include plant macroremains and other remains found in sediment samples) as indicators for preservation quality. Two late Neolithic lakeshore sites located in Central Switzerland were used as tests and more than 100 samples per site were investigated. Samples were grouped into meaningful groups (according to sediment type or their location in the stratigraphy, etc.) and ubiquities were calculated for each variable in each group of samples. Correspondence Analysis was applied in order to establish connections between groups of variables and groups of samples. GIS was used in one of the cases to look at preservation at a site scale. The method proved to be useful and differences in preservation conditions were observed in both sites, not only regarding the location of the samples in the stratigraphy and in relation to their proximity to the lake, but also in connection to sediment type. It is suggested that such studies are necessary before any palaeoeconomic analysis is undertaken.

Foliar nitrogen and phosphorus stoichiometry of three wetland plants distributed along an elevation gradient in Dongting Lake, China

We examined foliar nitrogen (N) and phosphorus (P) stoichiometry of 3 wetland plants (Phalaris arundinacea, Miscanthus sacchariflorus, and Carex brevicuspis) distributed along an elevation gradient in the Dongting Lake, China, and how this stoichiometry is related to soil physico-chemical characteristics, elevation, and flooding days. Plant and soil samples were collected from 3 lakeshore sites. Total N and P concentrations of plants and six physico-chemical characteristics of the soil were measured, in addition to the elevation and flooding days. P. arundinacea and M. sacchariflorus had higher total N and P concentrations than C. brevicuspis. The foliar N:P ratio decreased with increasing elevation, and only increased with increasing foliar total N concentration. Canonical correspondence analysis indicated that the foliar stoichiometry was primarily regulated by soil water content, followed by soil nutrient concentration. The foliar N and P stoichiometry of the 3 wetland plants was insignificantly correlated with soil total P concentration. However, foliar stoichiometric characteristics and soil total N concentration significantly differed among the 3 species. These results demonstrate that spatial variation of foliar stoichiometry in wetland plants exists along an elevation gradient, with this information being useful for the conservation and management of wetland plants in this lake.

Layers rich in aquatic and wetland plants within complex anthropogenic stratigraphies and their contribution to disentangling taphonomic processes

There is an ongoing discussion about how organic material is preserved in settlement layers on lakeshores. Different scenarios have been suggested; was a permanent water cover needed at all times, or were there episodical water level fluctuations? In this paper, we use aquatic and wetland plants to shed light on layer formation processes of complex anthropogenic stratigraphies of the Neolithic lakeshore settlement Zug-Riedmatt (Central Switzerland). Botanical macroremains from the microarchaeologically investigated occupation layer were analysed and compared to modern analogue data from two lakes. Results suggest that the excavated area of the settlement was in a first phase located in the sublittoral zone (below water), with a massive input of anthropogenic waste material contributing to the layer formation, but then in a second phase, the shoreline shifted so that the excavated part of the settlement was located in the eulittoral zone and in a third phase probably even landwards of a reed belt. In a comparison with two previously investigated Neolithic lakeshore sites, we found that at least in one of these sites, such ‘drier’ conditions could also be found. This shows how important it is to know which part of the littoral zone is represented in an excavation, as conditions can differ within a settlement, and that there is very likely no universally valid ‘Pfahlbau scenario’.