Statement of the problem. The hypothesis of this research is that it is possible to use the eye structure of aquatic animals, which performs well in low light conditions, to improve, optimize and introduce new daylighting systems. The intended transparent dome in this study is a double glass skin dome with placed modules between the two layers of dome to control daylighting. According to the natural structure investigated in this research, these modules are dynamic and variable in two fully open condition to receive maximum daylight in low dim light conditions (e.g., in the cloudy sky) and semi-closed in the completely sunny sky with intense radiation. Results and conclusions. What is intended as an independent variable in this research are: the opening width, and the shape of modules (square, hexagonal and triangular), the height of the modules and the rotation angle of the light reflecting panels of the modules. The dependent variables (appropriate illuminance and appropriate lighting distribution pattern) are investigated by Useful Daylight Illuminance (UDI). Radiance software have been used as daylight simulators. The results of this research include the optimal physical characteristics of the modules and the performance of variables on the illumination under the dome.