Abstract Ordovician Japan formed a mature arc-trench system developed along the palaeo-Pacific (Panthalassa) margin of the Greater South China (GSC) continental block. GSC consists of South China, East China Sea, SW–NE Japan and the Khanka–Jiamusi–Bureya megablock in the Far East; Paleozoic GSC was thus, in total, twice as large as the South China components by themselves (Yangtze and Cathaysia). The Ordovician crust of Proto-Japan comprised coeval arc-related rocks, such as granitoids, supra-subduction zone ophiolites and fore-arc basin strata, although most of them were considerably fragmented. The Ordovician and middle–late Paleozoic fossils from Japan are highly limited but suggest that Proto-Japan was positioned in the low-latitude domains probably of the palaeo-Pacific Ocean in connection to Paleo-Tethys. GSC became separated from Rodinia in the Neoproterozoic, and its Proto-Japan segment evolved as a collision-free subduction margin for nearly 500 myr since the mid-Cambrian. The GSC framework provides critical constraints to the palaeogeographical reconstruction of circum-Pacific continental blocks. First, the Cambro-Ordovician GSC should have been isolated from Australia/India/East Antarctica that formed East Gondwana by a relatively wide ocean domain for keeping ‘subduction potential’. Second, the Cathaysian margin of GSC should have faced to an extensive ocean without major continents since the Cambrian. The palaeo-Pacific is the only possible candidate for this.