Grass Suppression Research Articles

Hot Foam and Nitrogen Application to Promote Spring Transition of “Diamond” Zoysiagrass (Zoysia matrella (L.) Merr.) Overseeded with Perennial Ryegrass (Lolium perenne L.)

In transition areas, cool season turfgrasses are overseeded in autumn to maintain the high quality of dormant warm season turfgrasses, while in spring several agronomic methods (scalping, coring, topdressing, verticutting, irrigation, and targeted fertilization) or chemical desiccation are adopted to remove the cool season turfgrasses from the stand. To reduce chemical applications, several methods of “thermal weeding” have been experimented with, but little is known about these methods in zoysiagrass (Zoysia spp. Willd) spring transition. A study was conducted at the University of Pisa, Italy, on Manila grass (Zoysia matrella (L.) Merr., cv “Diamond”) (Zm) overseeded with perennial ryegrass (Lolium perenne L.) (Lp) with the aim of comparing different methods of cool season grass suppression (scalping and hot foam) and different application rates of nitrogen. To assess treatment effect, green cover, turf quality, turf color, shoot density, and some vegetation indices (GLI, DGCI and NDVI) were determined. An average green cover of at least 90% was obtained on all plots seven weeks after the treatments. While scalping had minor effects on turf appearance and on polystand composition, hot foam had a stronger effect on turf color, green cover, and turf quality in the weeks following application. Once it had recovered from the hot foam treatments, the turf had a greater number of Zm shoots and a relevant reduction of Lp shoots. The hot foam was very effective in suppressing Lp while maintaining Zm recovery capacity.

Utilizing cover crops for weed suppression within buffer areas of 2,4-D-resistant soybean

AbstractCover crops can be utilized to suppress weeds via direct competition for sunlight, water, and soil nutrients. Research was conducted to determine if cover crops can be used in label-mandated buffer areas in 2,4-D-resistant soybean cropping systems. Delaying termination of cover crops containing cereal rye to at or after soybean planting resulted in a 25 to more than 200 percentage point increase in cover crop biomass compared to a control treatment. Cover crops generally improved horseweed control when 2,4-D was not used. Cover crops reduced grass densities up to 54% at four of six site-years when termination was delayed to after soybean planting. Cover crops did not reduce giant ragweed densities. Cover crops reduced waterhemp densities by up to 45%. Cover crops terminated at or after planting were beneficial within buffer areas for control of grasses and waterhemp, but not giant ragweed. Yield reductions of 14% to 41% occurred when cover crop termination was delayed to after soybean planting at three of six site-years. Terminating the cover crops at planting time provided suppression of grasses and waterhemp within buffer areas and had similar yield to the highest-yielding treatment in five out of six site-years.

Effects of herbicide underdoses on the vegetative development of Panicum maximum cultivars

The advance of intercropping has generated the need for studies that evaluate methods of suppression of forage grasses in order to not harm the development of the commercial culture. The aim of this study is to evaluate the sensitivity of forage grasses to post-emergence herbicide application. To this end, an experiment was carried out in a greenhouse using a randomized complete block design in a 4 x 9 factorial scheme with four replicates. The first factor consisted of the forage grasses Urochloa ruziziensis, Panicum maximum cv. BRS Tamani, P. maximum cv. BRS Quenia, and P. maximum cv. BRS Zuri. The second factor consisted of the combination of eight herbicide treatments applied at post-emergence in association with atrazine (1200 g a.i. ha -1 ), besides a control treatment, without application. P. maximum cv. BRS Kenya was the least sensitive to post-emergence herbicide application. Tembotrione (42 and 84 g a.i. ha -1 ) and mesotrione (48 and 96 g a.i. ha -1 ) have potential for suppression of U. ruziziensis . Tembotrione (84 g a.i. ha -1 ), glyphosate (200 g a.i. ha -1 ), and nicosulfuron (7.8 g a.i. ha -1 ) have potential for suppression of the cv. BRS Kenya, and tembotrione (42 g a.i. ha -1 ) and mesotrione (96 g a.i. ha -1 ) have potential for the suppression of BRS Tamani. The P. maximum cv. BRS Zuri presented a higher plant height, shoot dry matter, and root dry matter than the other forages evaluated. For this forage, the use of nicosulfuron herbicides at post-emergence, regardless of the dose, reduced the shoot dry matter by 70%.

The effects of land use and microsite availability on early seedling recruitment of Acacia tortilis (synonym: Vachelia tortilis) in a southern African savanna

Many tree recruitment studies in African savannas target individual factors that limit recruitment, yet recruitment of some species could be limited by an interplay of environmental filters. This study investigated the effects of land use and microsite availability on early seedling recruitment of Acacia tortilis in riverine woodlands in north-western Zimbabwe. The riverine woodlands were divided according to the level of human disturbance into protected and unprotected sites. Each site was further divided into riverine and off riverine microsites. In each microsite, we laid five main plots (100 × 50 m) and nine seedling subplots (5 × 5 m) nested in each main plot, making a total of 60 main plots and 180 subplots. We found higher seedling recruitment in unprotected than protected sites. Although seeder density was twice greater in protected than unprotected sites, seed bank density was lower in the former by a factor of 1.5. Tree cover and herbaceous biomass was 1.4 and 1.8 times greater in protected than unprotected sites, respectively but were negatively correlated with seedling density. Across all microsites, herbivory-related mortality was less than 40% while drought-related mortality was more than 60%. Put together, the results suggest that higher recruitment in unprotected sites was most likely facilitated by domestic ungulates through seed dispersal and grass suppression which enhanced the quality of regeneration sites. We conclude that encroachment of A. tortilis in communal grazing sites is not only important for its conservation but for a variety of goods and services derived from it.

Efficiency of Cardboard Crowning on the Suppression of Invasive Alien Grasses

Invasive exotic grasses are among the biggest obstacles to the success of reforestation projects. The aim of this study was to evaluate the efficiency of the crowning with cardboard in suppressing the grasses Urochloa humidicola and Megathyrsus maximus aiming its use in reforestation areas. A field experiment was installed in a completely randomized design with two treatments consisted of crowning with cardboard and without crowning (control). The crowning with cardboard caused a progressive reduction of the biomass and was effective in causing senescence of both grasses in 80% or more of the crowning area between 20 and 50 days after the crowning (F test; p<0,05). These results demonstrate the effectiveness and potential of using the crowning with cardboard to control alien grass species in reforestation areas. This method brings as an innovation the possibility of placing cardboard directly on the grass after mowing, allowing greater simplicity and lower operating cost.

Long-Term Seeding Outcomes in Slash Piles and Skid Trails after Conifer Removal

Conifer removal in interior woodland ecosystems of the western US is a common management treatment used to decrease fire hazard and shift woodlands to more historical states. Woody material is frequently removed by skidding material off site and via slash pile burning. Assessing the long-term outcomes of seeding treatments after such ground disturbing activities is critical for informing future management and treatment strategies. Using two designed experiments from a central Oregon juniper woodland, we resampled slash piles and skid trails 8 years after seeding. Our objectives were to assess the long-term vegetation response to conifer removal, ground disturbance, and seeding source (cultivar and local) in slash piles and skid trails. We found that seeded species persisted in the long term, but abundance patterns depended on the species, seed source, and the type of disturbance. In general, there were more robust patterns of persistence after pile burning compared to skid trails. Seeding also suppressed exotic grass cover in the long term, particularly for the local seed source. However, the invasion levels we report are still problematic and may have impacts on biodiversity, forage and fire behavior. Our short-term results were not predictive of longer-term outcomes, but short- and long-term patterns were somewhat predictable based on species life history traits and ecological succession. The use of a mix of species with different life history traits may contribute to seeding success in terms of exotic grass suppression. Lastly, our results suggest that locally adapted seed sources may perform as well or better compared to cultivars. However, more aggressive weed treatments before and after conifer removal activities and wider seeding application may be needed to effectively treat exotic grass populations.

Effects of elevated CO2 on competition between native and invasive grasses.

Elevated atmospheric CO2 concentration increases the performance of invasive plants relative to natives when grown in monoculture, but it is unclear how that will affect the relative competitive abilities per se of invasive and native grasses grown together. We tested competitive outcomes for four native and four invasive perennial C3 and C4 grasses under ambient (390ppm) and elevated (700 or 1000ppm) CO2 concentrations in the greenhouse with non-limiting water and nutrients. We predicted that elevated CO2 would increase the competitive suppression of native grasses by invasive grasses. To test this, we determined the relative interaction intensity of biomass allocation for natives grown alone vs. those grown in native-invasive species pairs. We also measured photosynthetic traits that contribute to plant invasiveness and may be affected by elevated CO2 concentrations for species pairs in mixture to determine native-invasive relative performance. We found no effect of CO2 for the aboveground biomass and tiller production measures of interaction intensity or for relative performance for most of the measured photosynthetic traits. In competition, the invaders nearly always outperform natives in biomass and tiller production, regardless of CO2 level. The results suggest that increasing CO2 concentration alone has little effect on grass competitive outcomes under controlled conditions.

Adzuki bean [Vigna angularis(Willd.) Ohwi &amp; Ohashi], oilseed radish (Raphanus sativusL.), and cereal rye (Secale cerealeL.) as living mulches with and without herbicides to control annual grasses in sweet corn (Zea maysL.)

Annual grasses are difficult to control in sweet corn in Canada due to the scarcity of registered herbicides with grass activity. In addition to the potential soil health benefits, over-seeding living mulches into the cropping system may help sweet corn growers improve annual grass control by increasing competitive ground cover. To test this hypothesis, trials were established at three locations in Ontario and Quebec, Canada, in 2008 and 2009. At each location, sweet corn was over-seeded at the 4–6 leaf stage with one of three living mulches alone or in combination with an herbicide. The living mulch/herbicide pairings were adzuki bean (linuron + S-metolachlor), cereal rye (saflufenacil), and oilseed radish (pendimethalin). All living mulch treatments were compared with an untreated control and an industry standard (S-metolachlor/atrazine). When sweet corn was over-seeded with living mulches alone, the most effective annual grass control was provided by the cereal rye. The least effective living mulch was adzuki bean, but the combination of adzuki bean plus a herbicide was the most effective for annual grass suppression. The final marketable yields in all living mulch treatments were always lower than the industry standard. In spite of effective annual grass control, reduced yields may make the adoption of the tested living mulch species less attractive to conventional sweet corn growers.

Native lagomorphs suppress grass establishment in a shrub-encroached, semiarid grassland.

Shrub encroachment into arid grasslands has been associated with reduced grass abundance, increased soil erosion, and local declines in biodiversity. Livestock overgrazing and the associated reduction of fine fuels has been a primary driver of shrub encroachment in the southwestern United States, but shrublands continue to persist despite livestock removal and grassland restoration efforts. We hypothesized that an herbivory feedback from native mammals may contribute to continued suppression of grasses after the removal of livestock. Our herbivore exclusion experiment in southeastern Arizona included five treatment levels and allowed access to native mammals based on their relative body size, separating the effects of rodents, lagomorphs, and mule deer. We included two control treatments and replicated each treatment 10 times (n = 50). We introduced uniform divisions of lawn sod (Cynodon dactylon) into each exclosure for 24‐hr periods prior to (n = 2) and following (n = 2) the monsoon rains and used motion‐activated cameras to document herbivore visitations. In the pre‐monsoon trials, treatments that allowed lagomorph access had less sod biomass relative to other treatments (p < 0.001), averaging 44% (SD 36%) and 29% (SD 45%) remaining biomass after the 24‐hr trial periods. Following the onset of monsoons, differences in remaining biomass among treatments disappeared. Desert cottontails (Sylvilagus audubonii) were detected more frequently than any of the other 11 herbivore species present at the site, accounting for 83% of detections during the pre‐monsoon trials. Significantly more (p < 0.001) desert cottontails were detected during the pre‐monsoon trials (2,077) compared to the post‐monsoon trials (174), which coincided with biomass removal from lagomorph accessible treatments. We conclude that desert cottontails are significant consumers of herbaceous vegetation in shrub‐encroached arid grasslands and they, along with other native herbivores, may act as a biotic feedback contributing to the competitive advantage and persistence of shrubs.

Suppression of prairie grasses due to excess magnesium in a portion of a restored prairie

ABSTRACT In June 2002, the Department of Energy (DOE) began establishing the 60 ha Howell Prairie at the DOE Weldon Spring Site (WSS). In one area, the clay base is different from the other soil (subarea 2C). Vegetation sampling was conducted on four permanent plots across the prairie beginning in 2008, and shows that three of the four plots have strong establishment of native prairie species including prairie grasses. The fourth plot (subarea 2C), where the soil is different, shows significantly less native grass cover and stunted vegetation compared to the other three plots. One hundred twenty-five soil samples were taken in 6 different months and in 6 different years across the entire prairie restoration. Across the prairie, potassium (K) and phosphorus (P) were not limiting. The pH, organic matter content (OM), and cation exchange capacity (CEC) did not show trends related to the vegetation issues in subarea 2C. Ordination of the ratio of magnesium (Mg):K shows that Mg is very high in subarea 2C, which has been demonstrated to suppress the growth of prairie grasses. Subarea 2C contains interstratified kaolinite-smectite clay which contributes Mg to soil. It is hypothesized that an inexpensive, nondestructive treatment (addition of K) could be applied to remediate this area.

Optimum Vegetation Conditions for Successful Establishment of Planted Eastern White Pine (Pinus strobus L.)

The 10th-growing season performance of planted eastern white pine (Pinus strobus L.) seedlings was evaluated in response to herbaceous and woody vegetation control treatments within a clearcut and two variants of the uniform shelterwood regeneration system (single vs. multiple future removal cuts). Herbaceous vegetation control involved the suppression of grasses, forbs, ferns and low shrubs for the first 2 or 4 growing seasons after planting. Deciduous woody vegetation control treatments, conducted in combination with the herbaceous treatments within a response-surface design, involved the permanent removal of all tall shrubs and deciduous trees at the time of planting, at the end of the 2nd or 5th growing seasons, or not at all. In general, the average size of planted pine was related positively to the duration of herbaceous vegetation control and negatively to delays in woody control. White pine weevil (Pissodes strobi Peck) altered these trends, reducing the height of pine on plots with little or no overtopping deciduous woody vegetation or mature tree cover. Where natural pine regeneration occurred on these plots, growth was similar but subordinate to the planted pine. Data from the three sites indicate that at least 60% of planted pine may be expected to reach an age-10 height target of 2.5 m when overtopping cover (residual overstory + regenerating deciduous) is managed at approximately 65% ± 10%, and total herbaceous cover is suppressed to levels not exceeding 50% in the first five years. On productive sites, this combination may be difficult to achieve in a clearcut, and requires fairly rigorous vegetation management in shelterwood regeneration systems. Currently, synthetic herbicides offer the only affordable and effective means of achieving such vegetation control.

Stocktype and grass suppression accelerate the restoration trajectory of Acacia koa in Hawaiian montane ecosystems

Restoring degraded mesic-montane forests represents a major challenge in maintaining functioning ecosystems throughout the tropics. A key example of this lies in Hawai'i, where restoring native koa (Acacia koa, A. Gray) forests are a top conservation and forestry priority because of the critical habitat and high-value timber products that they provide. Efforts to restore koa forests, however, are directly impeded by extensive, non- native kikuyu grass (Pennisetum clandestinum Hochst. ex Chiov.) swards occupying deforested montane landscapes. In this study, we implemented a combination of grass suppression and selection of koa seedling stocktypes to measure outplanting performance in a naturalized site on the island of Maui. Seedlings were grown in a nursery in two root container sizes (111 and 207 cm 3 ) and subsequently outplanted into grass-dominated plots that were either untreated or suppressed with a high-rate herbicide combination of imazapyr and glyphosate (1.7 kg a.i. ha -1 , respectively), 30 days prior to planting. Across all treatments, seedling survival was high ((95 %). Thirty months after planting, trees from the larger stocktype had significantly greater growth in height and root-collar diameter. Initial grass suppression resulted in trees that were 34 % taller with 66 % larger root-collar diameters after 30 months. Herbicide treated plots also had significantly higher leaf area indices (2.6 vs. 1.8 m 2 m -2 ), indicative of higher photosynthetic capacity and canopy closure. Grass suppression increased soil temperature along with soil moisture in the first year followed by a dramatic drop in moisture corresponding to large growth

Establishing Legumes in a Tall Fescue Sward

Tall fescue [Schedonorus arundinaceus (Schreb.) Dumort.] is a common grass species in the eastern half of the USA, but legumes grown with it could provide benefits. Obstacles to legume establishment in fescue pastures include disease, insect damage, and grass competition. Experiments were performed in 2010 and 2011 at Site 1 to test the efficacy of insect control, disease control, and two methods of grass suppression on seedling establishment of three legumes. The highest seedling density of red clover (Trifolium pretense L.) was obtained with glyphosate treatment regardless of pest control and for clipping without pest control. No overall or consistent benefit was found for white clover (Trifolium pretense L.) or trefoil (Lotus corniculatus L.) from seed and foliar insecticides or fungicides, or for grass suppression by either clipping or glyphosate application. Experiments were performed in 2012 and 2013 at Site 2 to test the efficacy of grass suppression by grazing cattle or by clethodim application. Seedling density of red clover was not significantly improved by either treatment, but the density of birdsfoot trefoil was increased by clethodim, and the density of white clover was increased in 2013 by both methods of grass suppression. Overall, red clover was least sensitive to grass competition. Birdsfoot trefoil was better served by the immediate effect of chemical suppression, whereas white clover benefitted most from the lengthier grass suppression provided by grazing.

Is truffle brûlé a case of perturbational niche construction?

Aim of study: in the context of niche construction theory, the investigation was aimed at assessing if truffle brûlé is a case of niche construction by testing if the disappearance of grasses in the brûlé induces a non-arbitrary perturbation in the soil physical environment and, in that case, which are the physical processes involved.Area of study: a natural truffle bed located in the Italian Apennines inside an experimental truffle-producing area of the University of Perugia.Material and methods: Three aggregate size classes in the soil of a brûlé and of the neighbouring grass-covered area were determined with the wet-sieving technique in accordance with international standards. In the first part of the investigation, the area was sampled according to a systematic sampling design and the spatial patterns of aggregate size classes in the brûlé and in the neighbouring grass-covered area were compared by means of geostatistics. In the second part, the suppression of grasses in the brûlé was mimicked in the laboratory by removing roots from a sample collected in the grass-covered area and the effect of freeze-thaw cycles was tested by comparing a control treatment to other four treatments consisting in sub-samples equilibrated at a water tension of -2.5, -5, -10 and -20 kPa. Samples were submitted to a number of freeze-thaw cycles equal to those recorded in the area, analysed for aggregate size distribution and compared with univariate ANOVA.Main results: The aggregate size classes larger than 0.25mm displayed a spatial pattern comparable to that of the brûlé, with sharp changes along the boundaries of the brûlé itself. The laboratory experiment showed that such changes are attributable to freeze-thaw cycles that in one winter season may produce a significant decrease in aggregate size compared to the grass-covered area. Both results indicate that the disappearance of grasses in the brûlé fulfils the requirement of perturbational niche construction.Research highlights: the observed change of soil aggregation in the brûlé can be useful in delineating soils suitable for truffle farming and in creating suitable soil physical conditions for truffle production by exploiting freeze-thaw cycles after irrigation in winter.Keywords: aggregates; brûlé; niche construction; Tuber, soil structure.

Intercropping of soybean cultivars with Urochloa

The integrated crop-livestock system is a key strategy for both grain and livestock production. The cultivation of grass species in between two soybean cropping seasons has been an important component of this system, in Brazil. Thus, this study aimed to evaluate the yield of glyphosate-resistant soybean cultivars intercropped with Urochloa species, under different forage management systems. A total of six experiments, consisting of three soybean cultivars (BRS 295 RR, BRS 316 RR, and BRS 294 RR) and two grass species (Urochloa ruziziensis and Urochloa brizantha), were established. For each experiment, four treatments (soybean only; grass only; soybean intercropped with a grass species, with the application of glyphosate to decrease the growth of grass; and soybean intercropped with a grass species, without growth suppression) were evaluated in a completely randomized experimental design, with four replications. For each experiment, only one soybean cultivar and one grass species were used. The grasses were sown in the soybean inter-rows at 27 days after sowing (DAS), while the glyphosate application to the treatment with grass suppression was performed at 84 DAS for the soybean or 57 DAS for the grasses. The yields of the three soybean cultivars were not significantly affected by either grass species, even though the shoot dry-matter yields for both of the intercropped grass species were high, though lower than the values obtained without the interference imposed by soybean.

Overcoming barriers to seedling regeneration during forest restoration on tropical pasture land and the potential value of woody weeds.

Combating the legacy of deforestation on tropical biodiversity requires the conversion to forest of large areas of established pasture, where barriers to native plant regeneration include competition with pasture grasses and poor propagule supply (seed availability). In addition, initial woody plants that colonise pasture are often invasive, non-native species whose ecological roles and management in the context of forest regeneration are contested. In a restoration experiment at two 0.64 ha sites we quantified the response of native woody vegetation recruitment to (1) release from competition with introduced pasture grasses, and (2) local facilitation of frugivore-assisted seed dispersal provided by scattered woody plants and artificial bird perches. Herbicide pasture grass suppression during 20 months caused a significant but modest increase in density of native woody seedlings, together with abundant co-recruitment of the prominent non-native pioneer wild tobacco (Solanum mauritianum). Recruitment of native species was further enhanced by local structure in herbicide-treated areas, being consistently greater under live trees and dead non-native shrubs (herbicide-treated) than in open areas, and intermediate under bird perches. Native seedling recruitment comprised 28 species across 0.25 ha sampled but was dominated by two rainforest pioneers (Homalanthus novoguineensis, Polyscias murrayi). These early results are consistent with the expected increase in woody vegetation recruitment in response to release from competitive and dispersive barriers to rainforest regeneration. The findings highlight the need for a pragmatic consideration of the ecological roles of woody weeds and the potential roles of “new forests” more broadly in accelerating succession of humid tropical forest across large areas of retired agricultural land.

Selection of crop cultivars suited to the location combined with astute management can reduce crop yield penalties in pasture cropping systems

Pasture cropping is an emerging farming-systems practice of southern Australia, in which winter grain crops are sown into an established stand of a winter-dormant, summer-growing perennial pasture. There is a pressing need to define times, locations and climates that are suitable for pasture cropping. To evaluate effects of management interventions, agro-environment, and possible interactions on crop and pasture productivity associated with pasture cropping, an AusFarm® simulation model was built to describe a pasture-cropping system based on annual crop and subtropical grass. The model was parameterised using data from field research on pasture cropping with barley cv. Buloke and a C4 subtropical grass, Gatton panic (Panicum maximum cv. Gatton), conducted at Moora, Western Australia. The simulation was run over 50 years using the historical climate data of five southern Australian locations (Cunderdin, Jerdacuttup, Mingenew, and Moora in Western Australia, and Karoonda in South Australia). Two wheat cultivars and one barley crop were considered for each location, to examine the impact of crop phenology on this farming system. Jerdacuttup and Moora favoured pasture cropping, with average barley-yield penalties of 10 and 12%. These locations were characterised by colder growing seasons, more plant-available water at anthesis, and more winter–spring rain. The cereal crops did not rely on stored soil moisture, growing instead on incident rain. The winter–spring growth of the Gatton panic pasture was highest at Mingenew. This generated a high yield penalty, 38% loss under pasture cropping, compared with the other locations. Changing the efficacy of a herbicide application to the pasture when the crop was sown had a strong effect on yield. Yield penalties at Moora and Mingenew reduced to 7 and 29%, respectively, when the proportion of live biomass killed by the herbicide was doubled. Utilisation of soil moisture by the Gatton panic pasture during summer and early autumn had little effect on subsequent grain yield, whereas reduced pasture growth during the winter–spring growing period had a substantial effect on crop yield. Pasture cropping can therefore succeed in agro-climatic regions where crops can be grown on incident rain and pasture growth is suppressed through low temperature or herbicide. Perennial pasture growth should be minimised during the crop growing period through the management of crop sowing date, nitrogen fertiliser application and C4 grass suppression to minimise the effect on stored soil water at crop anthesis.

Declining diversity in abandoned grasslands of the carpathian mountains: do dominant species matter?

Traditional haymaking has created exceptionally high levels of plant species diversity in semi-natural grasslands of the Carpathian Mountains (Romania), the maintenance of which is jeopardized by recent abandonment and subsequent vegetation succession. We tested the hypothesis that the different life history strategies of dominant grasses cause different patterns of diversity loss after abandonment of traditional haymaking in two types of meadow. Although diversity loss rate was not significantly different, the mechanism of loss depended on the life history of dominant species. In meadows co-dominated by competitive stress-tolerant ruderals, diversity loss occurred following the suppression of dominant grasses by tall forbs, whereas in meadows dominated by a stress-tolerant competitor, diversity loss resulted from increased abundance and biomass of the dominant grass. We conclude that management for species conservation in abandoned grasslands should manipulate the functional turnover in communities where the dominant species is a weaker competitor, and abundance and biomass of dominant species in communities where the dominant species is the stronger competitor.

Restoration of native plant communities in a Hawaiian dry lowland ecosystem dominated by the invasive grass Megathyrsus maximus

AbstractQuestionsHow does a highly degraded Hawaiian tropical dry lowland ecosystem dominated by the non‐native invasive Megathyrsus maximus (guinea grass) respond to different restoration treatments (three native species outplanting treatments; four native broadcast seed treatments)? What effect do restoration treatments have on invasive and native species groundcover, biomass and physiological activity, and volumetric soil water content?LocationWaianae Kai Forest Reserve, Island of Oahu, Hawaii, USA.MethodsThe invasive grass M. maximus was suppressed by initial mowing and pre‐ and post‐planting herbicide applications. Native species were added in three outplant and four broadcast seed treatments in a complete randomized block design. Native species and M. maximus growth and ecophysiology, and volumetric soil water content were quantified for 8 mo following treatment establishment.ResultsNative species outplant survival ranged from 38% to 67%. Cover of M. maximus was significantly reduced in all outplant treatments compared with control and treated control (mowing and herbicide without native species additions), but did not differ across outplant treatments. Of the native species, Dodonaea viscosa biomass was higher than Cordia subcordata, while other native species did not differ. Maximum photosynthetic rates (Amax) did not differ across species in July. However, in August (drier period), M. maximus exhibited lower Amax than all native species except T. populnea, indicating adaptive dormancy during drought. Broadcast seeding with native species was not an effective restoration treatment, as field germination ranged from 0.5% to 2.3%.ConclusionsEcological restoration of highly invaded Hawaiian tropical dry lowland ecosystems can be mediated through aggressive invasive species suppression and native species outplanting. Recommendations for restoration include initial removal of invasive grasses, adaptive suppression of grasses post‐outplanting, and utilization of diverse native species assemblages that are ecophysiologically adapted to local conditions and competitive with M. maximus.

Influence of herbaceous and woody competition on white pine regeneration in a uniform shelterwood

We investigated the effects of herbaceous and woody vegetation control on the survival and growth of planted and natural eastern white pine (Pinus strobus L.) seedlings through six growing seasons following a uniform shelterwood regeneration harvest on two independent sites. Subsequent to chain scarification, white pine seedlings were planted at 2-m spacing, augmenting natural regeneration (full stocking and &gt;3000 seedlings per ha). Herbaceous vegetation control involved the suppression of grasses, forbs, ferns, and low shrubs, and was maintained for zero, two, or four growing seasons after planting. Woody control involved the removal of all tall shrubs and deciduous trees, and was conducted at the time of planting, at the end of the second or fifth growing seasons, or not at all. White pine seedling growth responded positively to increased duration of herbaceous vegetation control and negatively to delayed woody control. Maximum growth was not realized unless both types of vegetation were suppressed. During the first six growing seasons, the height growth of planted pine was more than twice that of naturally regenerating pine, regardless of tending regime. The study suggests that successful white pine regeneration may be achieved by thinning from below to allow 50% to 60% of full sunlight in the understory, followed by the proactive, early suppression of woody and herbaceous vegetation to maintain optimum light levels and reduce competition for soil moisture and nutrients.