Fossil Bivalves Research Articles

Evolution of left–right asymmetry in side-resting (pleurothetic) bivalves: a test of genes as leaders or followers

ABSTRACT Side-resting (pleurothetic) lifestyles evolved independently many times across the Bivalvia in free-lying, byssally attached and cementing lineages. Within some species individuals lie indifferently on either their right or left side (randomly pleurothetic), whereas in others they lie consistently on one side (left-pleurothetic or right-pleurothetic). Morphological differences between valves (asymmetries) typically accompany this behaviour, so it can be studied widely across living and fossil bivalves. Because direction of asymmetry is rarely inherited in cases of random asymmetry, any evolutionary transition from randomly pleurothetic to fixed (left or right) qualifies as a case of ‘genes as followers’. Origins of left-pleurothetic or right-pleurothetic taxa directly from upright (orthothetic) ancestors would represent examples of ‘genes as leaders’. I therefore surveyed pleurothetic lifestyles across the Bivalvia. By mapping pleurothetic states onto a comprehensive phylogenetic tree that included both living and fossil families, I assessed the prevalence of these alternate modes of evolution. Various lines of evidence were also used to hypothesize phylogenetic relations within the Chamidae—the only living bivalve family exhibiting all three pleurothetic states. Randomly pleurothetic forms (direction not inherited) arose from orthothetic ancestors about half as often as fixed pleurothetic forms (direction is inherited). Unlike many other animal groups, fixed pleurothetic forms rarely arose from randomly pleurothetic ancestors, suggesting a ‘genes as leaders’ mode of evolution was most prevalent. However, many fascinating examples of randomly pleurothetic forms among Upper Ordovician and Silurian bivalves suggest that evolutionary transitions from random to fixed may have been more common. Also, randomly pleurothetic forms were short-lived in some fossil lineages, which would reduce the likelihood of them being detected. Hopefully, this preliminary analysis will motivate further study of (1) phylogenetic relations of key groups like the Chamidae, and basal Limida, Ostreida and Pterioidea, and (2) morphological variation among early members of other pleurothetic bivalve groups.

Shallow-water temperature seasonality in the middle Cretaceous mid-latitude northwestern Pacific

Temperature seasonality during the middle Cretaceous provides vital information about climate dynamics and ecological traits of organisms under the conditions of the “supergreenhouse” Earth. However, sub-annual scale paleotemperature records in the mid-latitude region remain limited. In this study, sclerochronological and stable oxygen isotope (δ18O) analyses of bivalve fossils from the northwestern Pacific (paleolatitude: 44°N) were used to estimate their life history and sub-annual scale temperature patterns of the middle Cretaceous. The materials studied included Cucullaea (Idonearca) delicatostriata and Aphrodina pseudoplana recovered from middle Turonian (middle Cretaceous) shallow marine deposits in Hokkaido, northern Japan. Growth increment width and shell δ18O of C. (I.) delicatostriata revealed that the growth rate was temporally maximized and then minimized, which can be interpreted as representing spring and winter growth, respectively. Approximately 25 fortnightly growth increments occurred within that cycle, suggesting that shell formation proceeded continuously throughout the year. Based on shell δ18O values, shallow-water temperatures from 28°C to 35°C with 7°C seasonality were estimated, under the assumption that seawater δ18O values were annually invariant at −1‰ relative to VSMOW. This temperature seasonality in the middle Cretaceous is more than 5°C smaller than the seasonality of modern shallow-water environments at the same latitudes. These findings, taken together with previous studies of other oceanic regions, suggest that the Northern Hemisphere had low seasonal shallow-water temperature variation of up to 10°C in the middle Cretaceous.

Clumped isotope records of terrestrial temperatures during the Middle Jurassic (180–150 Ma) in East China

The Jurassic is a crucial greenhouse period that witnessed a transition from the relatively colder Triassic to the relatively warmer Cretaceous. However, previous studies based on marine sections suggest different long-term paleotemperature trends throughout the Jurassic. Oxygen isotope (ẟ18O) values of temperate belemnite and bivalve fossils suggest a general cooling followed by a general warming trend during the Jurassic, whereas TEX86H and clumped isotope sea surface temperature records available for parts of this period appear consistent with a relatively warm and stable climate. The paucity of Middle Jurassic paleotemperature estimates limits efforts to investigate the above disagreement. Here, we apply clumped isotope (Δ47) thermometry to paleosol carbonates from the central part of East China to reconstruct terrestrial paleotemperatures and paleoatmospheric CO2 (pCO2) in the Middle Jurassic. After evaluating potential burial diagenesis and seasonality of soil carbonate accumulation, we suggest that terrestrial mean annual temperatures ranged from 30 ± 4 °C to 34 ± 4 °C and the estimated mean pCO2 was 1219 ± 519 ppmv in mid-latitude East China during the Middle Jurassic. Based on the compiled marine temperatures and our results, we conclude that the paleoclimate was relatively warm and stable from the Middle Jurassic to at least the Earliest Cretaceous (ca. 180–135 Ma). Our estimated Middle Jurassic pCO2 aligns with previous records, revealing minimal variations compared to the pCO2 observed during the Early Cretaceous. These findings imply a link between the consistent temperatures and the relatively stable pCO2 levels from the Middle Jurassic through the Early Cretaceous.

Palynological investigation of an Upper Triassic sequence in Nanjing area, eastern China, with paleoecological implications

The end-Triassic is characterized by a significant biotic crisis in both marine and terrestrial realms. However, terrestrial records in eastern Tethys are relatively limited, especially in eastern China. The Fanjiatang Formation represents the Upper Triassic sequence in southern Jiangsu and Anhui provinces of eastern China, yielding a rich plant and bivalve fossil record. The present study provides the first detailed palynological investigation for the Upper Triassic of Nanjing area, eastern China. Six of twenty processed palynological samples are productive, and more than 70 terrestrial spore and pollen fossil taxa were identified. A palynological assemblage, namely, the Concavisporites–Dictyophyllidites–Kyrtomisporis–Classopollis Assemblage was established, characterized by abundant fern spores, mainly including Concavisporites, Dictyophyllidites and Kyrtomisporis, and much less abundant gymnosperm pollen grains (important taxa include Alisporites, Chasmatosporites, and Classopollis), indicating a Late Triassic Norian–Rhaetian age. In conjunction with previously reported macro-plant data, the palynofloral analysis reveals a vegetation dominated by ferns, with diverse cycadophytes, less abundant lycophytes, bryophytes, equisetales, ginkgophytes and seed ferns, and a few conifers. Paleovegetation and semiquantitative palynological analysis permit a paleoecological interpretation, suggesting a lowland groundcover-dominated vegetation under (sub)tropical warm and humid climate in Nanjing area during this time interval. More detailed palynological and macro-plant data through the whole Upper Triassic in this region are necessary in the future to better reveal the Late Triassic paleoecological settings and variations on land prior to the end-Triassic mass extinction in eastern Tethys.

A taxonomic revision of Neodietrichia (Araneae: Linyphiidae), a rarely encountered but widespread spider taxon.

The linyphiid spider Dietrichia hesperia was described from California in 1933 by Crosby and Bishop. The genus was renamed to Neodietrichia in 2008 by Özdikmen since Dietrichia was already occupied by a fossil bivalve. Until now, this rarely collected arboreal spider was the sole member of Neodietrichia. Herein we revise the genus by redescribing Neodietrichia hesperia (Crosby and Bishop, 1933), transferring Lophomma depressum (Emerton, 1882) to Neodietrichia as Neodietrichia depressum n. comb., and provide the first description of the female of this species.

Temporal shell-size variations of bivalves in South China from the Late Permian to the early Middle Triassic

During the end-Permian mass extinction and subsequent Early-Middle Triassic recovery, a series of environmental changes occurred and were accompanied by morphological and ecological responses. This study presents a shell-size analysis study of Upper Permian through lower Middle Triassic bivalve fossils from South China to discover their ecological and environmental implications. With a dataset of over 4900 bivalve specimens from published monographs and our own collections, shell length and shell height are measured and analyzed with a substage-stage binned scale. The mean and median sizes of bivalve shells show a significant Lilliput effect in the immediate aftermath of the end-Permian mass extinction. However, both epifaunal and infaunal bivalves grew larger quickly during the late Griesbachian, indicating a quick rebound of bivalve shell size. The temporal variations in bivalves in South China were mainly controlled by biotic and abiotic factors, e.g., physiological adaptation, changing competition, predatory pressure, and climatic and oxygen changes.

Implications of Late Palaeozoic postglacial marine transgressive‐regressive (T‐R) cycles recorded in the Talchir Formation, Son Valley Basin, peninsular India: A sequence stratigraphic paradigm

The signature of Late palaeozoic glaciation, a pan‐Gondwanaland palaeoclimatic event, is recorded in the Talchir Formation (Late Carboniferous to Early Permian), the basal‐most litho‐unit within the Gondwana Supergroup (Late Carboniferous–Early Cretaceous) in peninsular India. About 200 m thick sedimentary succession of the Talchir Formation, exposed in the Hasdeo River section in the Son Valley Basin, Central India, manifests signatures of deposition by melting of glaciers near to the ice‐grounding line. Evidence of glacial influences is prevalent in the lower‐middle part of the succession and represented by glacial tillites with faceted/striated gravels and bullet‐shaped clasts, dropstones and ice‐rafted debris (IRD), associated with various sediment gravity flow deposits. In the middle‐upper part of the succession, records of post‐glacial marine incursions are preserved as thick sandstone and sandstone‐mudstone heteroliths of a shallow marine shore‐shelf depositional regime. These are characterized by abundant tidal and wave signatures, with marine bivalve fossils and trace fossils of marine invertebrates. The upward decrease in size and abundance of the IRDs indicates a gradual decrease in glacial influences, with landward incursion of the marine depositional systems over the proglacial systems. Multiple stacked progradational and retrogradational stratal stacking patterns formed by alternating glacigenic and marginal marine dominated processes signify multiple transgressive‐regressive (T‐R) cycles, manifesting repeated advancements and retreats of the glacier, associated with sea level rise and marine incursions into the land. The top part of the succession, characterized by fine‐grained siltstone/mudstone without any IRD, suggests sedimentation at a relatively deeper part of the basin, indicating the strong influence of the eustatic rise caused by significant melting of the retreating ice sheet. Stacked T‐R cycles bounded by distinct facies architecture define multiple system tracts confined within three major deglaciation sequences within the studied Talchir Formation, suggesting multiple phases of advance‐retreats of the ice sheet and significant marine onlaps within the continental Gondwanaland under the Late Palaeozoic icehouse‐greenhouse climatic change.

Early Late Triassic retro-foreland basin in response to flat subduction of the Paleo-Tethyan oceanic plate, SE Tibet

Syn-subduction basins bear significant implications to understand tectonic evolution of any fossil subduction zone. The late Paleozoic to early Mesozoic (Paleo-Tethyan) tectonics of the eastern and southeastern Tibetan Plateau (i.e., the Sanjiang Orogenic Belt) is featured by ocean-continent subduction systems. A huge pile of volcanic-absent sedimentary succession developed in the middle segment of the Sanjiang orogenic belt, its age and tectonic nature remain unclear. Detailed geological mapping and zircon U-Pb dating results demonstrate that the early Late Triassic volcanic-absent succession comprises the nonmarine Maichuqing Formation in the lower part and the shallow marine Sanhedong Formation in the upper part. The Maichuqing Formation consists of coarse to fine-grained sandstone, siltstone and mudstone with abundant basal erosional surfaces, trough and planar cross-beddings, ripples, mudcracks, and plant fragments. The Sanhedong Formation comprises predominantly bioclastic limestones interlayered with marl, calcareous-muddy siltstone, and calcareous sandstone with abundant bivalve fossils. Syn-sedimentation deformation structures, such as slump folds and associated normal faults are common, suggesting intense tectonism during deposition. Synthesizing sedimentary data, paleocurrent and provenance results, combined with other available data, demonstrate that the volcanic-absent succession deposited within a retro-foreland basin along the rear part of the Permian-Triassic Jomda-Weixi-Yunxian arc in response to flat-subduction of the Paleo-Tethyan Ocean during the early Late Triassic time.

The earliest example of sexual dimorphism in bivalves—evidence from the astartid Nicaniella (Lower Jurassic, southern Germany)

AbstractProtandrous sex change (sex change from male to female) is one of the diverse sexual expressions exhibited by bivalves, but its expression in the shell is quite rare. Previous studies on living and fossil astartids suggest a relationship between protandrous sex change and the formation of crenulations on the ventral shell margin at later ontogenetic stages. Here we report the formation of such crenulations in the Early Jurassic astartid Nicaniella rakoveci (Kuhn, 1935) from the Amaltheenton Formation at Buttenheim, Franconia. This is the earliest known record of protandrous hermaphroditism in fossil bivalves, predating previous reports by at least 13 Myr. A principal component analysis of linear size measurements and Fourier shape analysis of the shell outlines revealed that the outline of Nicaniella rakoveci specimens varies from subquadrate to subcircular, but this variation is independent of the presence or absence of crenulations and therefore not associated with sex. Crenulated specimens exhibit a lower height/inflation ratio than non-crenulated ones, suggesting that the protandrous females have more inflated valves than the males. The formation of crenulations was probably related to allocation of resources for reproduction. The most likely function of the crenulations was to increase the internal shell volume in the female stage to accommodate more eggs rather than being an adaptation against predation as often assumed for other bivalves. The formation of crenulations is part of the protandrous life history and probably is controlled by a genetic mechanism that is also responsible for sex change.

Fossil bivalves and the sclerochronological reawakening

AbstractThe field of sclerochronology has long been known to paleobiologists. Yet, despite the central role of growth rate, age, and body size in questions related to macroevolution and evolutionary ecology, these types of studies and the data they produce have received only episodic attention from paleobiologists since the field's inception in the 1960s. It is time to reconsider their potential. Not only can sclerochronological data help to address long-standing questions in paleobiology, but they can also bring to light new questions that would otherwise have been impossible to address. For example, growth rate and life-span data, the very data afforded by chronological growth increments, are essential to answer questions related not only to heterochrony and hence evolutionary mechanisms, but also to body size and organism energetics across the Phanerozoic. While numerous fossil organisms have accretionary skeletons, bivalves offer perhaps one of the most tangible and intriguing pathways forward, because they exhibit clear, typically annual, growth increments and they include some of the longest-lived, non-colonial animals on the planet. In addition to their longevity, modern bivalves also show a latitudinal gradient of increasing life span and decreasing growth rate with latitude that might be related to the latitudinal diversity gradient. Is this a recently developed phenomenon or has it characterized much of the group's history? When and how did extreme longevity evolve in the Bivalvia? What insights can the growth increments of fossil bivalves provide about hypotheses for energetics through time? In spite of the relative ease with which the tools of sclerochronology can be applied to these questions, paleobiologists have been slow to adopt sclerochronological approaches. Here, we lay out an argument and the methods for a path forward in paleobiology that uses sclerochronology to answer some of our most pressing questions.

Correlation between the Warepan/Otapirian and the Norian/Rhaetian stage boundary: implications of a global negative δ13Corg perturbation

ABSTRACT The Norian/Rhaetian boundary interval (Late Triassic) is characterised by significant biotic turnover and severe climatic changes. However, this boundary still awaits formal definition. We have investigated the δ13Corg record within a well-preserved marine sequence exposed in the Kiritehere section, North Island, New Zealand. This section was located at higher latitudes (above 60°S) during the Triassic, and is considered the best exposed section representative of the New Zealand Warepan/Otapirian stage boundary. This is a regional boundary pertaining to the Zealandia region of Gondwana; it is described in the New Zealand Geological Timescale (NZGT) and it has been recognised as being correlative with the Norian/Rhaetian boundary. Using an integrated chemo- and biostratigraphic approach, we have identified a negative δ13Corg shift close to the disappearance of the standard-sized monotids (fossil bivalves) in the Kiritehere section. We interpret this negative shift to be the same negative δ13Corg shift that has been proposed to define the base of the Rhaetian. However, the position of the negative δ13Corg shift in the Kiritehere section is ca. 6.40 m below the accepted position of the Warepan/Otapirian boundary, based on biostratigraphy. This δ13Corg isotopic shift, best thought of as a chemostratigraphic event, is regarded here as a global phenomenon.

Performance of 3D Morphological Methods in the Machine Learning Assisted Classification of Closely Related Fossil Bivalve Species of the Genus Dreissena

In recent years, 3D analyses, new indices to describe the complexity of morphological structures and sophisticated machine learning approaches have advanced morphometrical analyses to assist species determination. However, the applicability of these modern approaches to the determination of cryptic species or fossil taxa has rarely been investigated.In this study, fossil and subfossil specimens of the four modern Dreissena species in the Caspian Sea are used to test the performance of 3D-based morphological approaches for machine learning assisted species identification. Specifically, 3D scans of the shells were used to construct 3D models for calculating “traditional” shell dimensions and “modern” shell complexity parameters. Finally, two machine learning approaches were applied to test the determination performance of shell measurements vs. shell complexity and individual vs. combined shell parameters.The results show that (i) there is no superior machine learning approach to species determination based on shell characters, (ii) shell complexity parameters are not per se more suitable for species identification than shell dimensions, (iii) a combination of shell parameters increases determination performance and reduces their species dependence and (iv) shell characters alone do not allow precise determination of all Dreissena species studied.These findings suggest that the most appropriate machine learning approach, the most informative shell characters and the best combination of characters need to be tested individually for different data sets. However, considering that it is difficult even for experts to distinguish Dreissena species based on shell characters, the machine learning assisted classification in the current study has performed comparatively well. Future analyses based on machine learning may therefore help experts to process large sample sizes efficiently and non-specialists to assess species level information with reasonable certainty.

The Discovery of the Bivalve Fossil Arguniella from the Lower Cretaceous Shangkuli Formation in Northern Greater Khingan Mountains, China: Implications for Stratigraphic Correlation

The Discovery of the Bivalve Fossil <i>Arguniella</i> from the Lower Cretaceous Shangkuli Formation in Northern Greater Khingan Mountains, China: Implications for Stratigraphic Correlation

New freshwater mussels (Bivalvia, Unionida) with potential trigonioidid and hyriid affinities from the Early Cretaceous of Brazil

Two new taxa of freshwater mussels (Bivalvia: Unionida) from the Aptian Crato Formation of the Araripe Basin, NE Brazil, are described. The fossil bivalves are confined to 30- to 130-cm-thick bioturbated mudstones overlying the fossil-rich laminated limestones of the Crato Formation Konservat-Lagerstätte. Individuals are often preserved with closed or splayed articulated valves, some of them potentially in life position, forming an autochthonous to parautochthonous assemblage. Monginellopsis bellaradiata nov. gen., nov. sp. shares key characters with the Trigonioidoidea: (i) the anterior pedal retractor muscle scar is clearly separated from the anterior adductor muscle scar; (ii) the shell has fold-like radial ribs on the posterior half; (iii) a right valve anterior tooth has a striated facet. Araripenaia elliptica nov. gen., nov. sp. is the most abundant and widely distributed unionid of the Crato Formation. Its ornament of anterior inverted V-shaped riblets, and central and posterior radial and sub-radial riblets resembles modern and fossil Hyriidae from the Americas, but also Trigonioidoidea from Eurasia. Its dentition of two smooth anterior pseudocardinals and two smooth posterior laterals in each valve provides no further clues for systematic assignment; muscle scars are not preserved. Assignment to the Hyriidae would make Araripenaia the oldest member of this family known from South America. Moreover, this bivalve assemblage of trigonioidoidids, hyriids, and previously reported silesunionoids suggests palaeobiogeographic links to other areas in both Gondwana and Laurasia.

Strontium stratigraphy of the Oligocene–Early Miocene shellbeds of the Kutch Basin, western India, and its implications

The Kutch Basin is unique among the western Indian sedimentary basins because of its near-complete sequence of post-Palaeozoic rocks. Due to extensive marine influence, the Oligocene–Early Miocene formations of the basin, namely Maniyara Fort, Khari Nadi and Chhasra, contain numerous shellbeds. Although age assignments of these formations exist based on foraminiferal biostratigraphy, detailed numerical age of the lithounits are yet to be established. We have identified a total of eleven distinct shellbeds (oldest SB 01 to youngest SB11) from this interval primarily containing bivalve fossils. Using 87Sr/86Sr of selected oyster and pectinid shells with pristine shell characteristics, we report the age of four shellbeds. The ages of SB 01, SB 04, SB 06 and SB 10 are 24.37, 17.31, 16.85 and 15.38 Ma, respectively. Our dates suggest a Chattian (24.37 Ma) age for SB 01 from the Bermoti Member, validating the previous biostratigraphical estimates from the Maniyara Fort Formation. The Chhasra Formation, however, shows a younger range of ages (17.31–15.38 Ma) characterized by a transition from the Burdigalian (SB 04–SB 06) to the Langhian (SB 10) stages. These dates have important implications in the study of sequence stratigraphy, Palaeobiogeography and tectonic history of the Kutch Basin. A surface with subaerial exposure is found in SB 08 (between 16.85 and 15.38 Ma) that corresponds to a global eustatic sea-level decrease (Mi2). Our new dates will also help evaluate the response of marine fauna to the closure of the Tethyan seaway around 19 Ma due to the formation of ‘Gomphotherium Landbridge’. The dated shellbeds enable us to identify pre- and post-closure fauna and assess the effect of biogeographical separation on these fauna. These dates have important implications in evaluating the regional geological record of western India in the context of various global events.

New Evidence for a Cretaceous Age for a Mesozoic Nonmarine Bivalve Assemblage from Paekto‐dong, Sinuiju City, The Democratic People's Republic of Korea

The Sinuiju Formation in Paekto‐dong, Sinuiju City in the Democratic People's Republic of Korea has yielded Mesozoic nonmarine bivalve fossils, which is the first occurrence of such in the DPRK. Based on these fossil specimens, a new Cretaceous bivalve assemblage, the Arguniella yanshanensis‐Sphaerium anderssoni Assemblage is erected. This assemblage includes Arguniella yanshanensis, A. lingyuanensis and Sphaerium anderssoni and can be compared with the Jehol Biota. The age of the Sinuiju Formation is also clarified and on the basis of the bivalves and the presence of a Eosestheria–Ephemeropsis–Lycoptera (E–E–L) assemblage, the formation is not Upper Jurassic, but Lower Cretaceous in age.

Drenov Grič black limestone: a heritage stone from Slovenia

Abstract Drenov Grič black limestone is considered to be one of the most beautiful Slovenian natural stones due to its black colour interwoven with white veins. Over the centuries, it has been extracted from two major quarries located west of Ljubljana. One of these quarries has been declared a valuable natural feature of national importance and is protected as a natural monument. This well-stratified, Triassic (Carnian) micritic limestone occurs in 10–80 cm thick beds with thin marl interlayers. The limestone occasionally contains abundant fossil bivalves, gastropods and ostracods. It is relatively rich in carbonaceous and bituminous organic matter, which is responsible for the black colour of the stone. The stone has been widely used in Slovenian monuments. Many indoor and outdoor architectural elements have been constructed using this limestone, particularly during the Baroque period, which was known for its extensive use of black limestones in other European countries as well. The most significant use of this limestone has been recorded in sculpted portals and altars. Some important buildings, which were decorated utilizing this stone, have been declared cultural monuments of local or national importance. Use of this limestone was also documented in other European countries (Italy, Austria, Serbia) and worldwide (USA). When exposed to climatic influences, chromatic and salt weathering are recognized as the main deterioration phenomena for this limestone when used in monuments.

Magnetostratigraphic dating of a drill core from the Northeast Plain of China: Implications for the evolution of Songnenpaleo-lake

Lacustrine sediments from the Northeast Plain of China are an ideal archive for recording the climatic history and variability of the East Asian monsoon during the Quaternary in mid-latitude East Asia. However, the lack of a high-resolution stratigraphy and high-precision geochronology of these sediments has limited our understanding of the mechanisms of Quaternary climatic and environmental changes in the region. In this study, we present a high-resolution magnetic susceptibility stratigraphy and precise paleomagnetic and optically stimulated luminescence (OSL) chronologies for core Qiananlingzi (QAL) from the Northeast Plain. Paleomagnetic data were obtained using alternating-field (AF) demagnetization with a peak AF of 80 mT and increments of 2.5−10 mT. In addition, OSL dating was conducted on coarse-grained quartz (90−150 µm) using the single aliquot regenerative dose (SAR) protocol. Correlation of the recognized polarity intervals of the QAL core to the geomagnetic polarity timescale was achieved by combining the magnetostratigraphic and OSL geochronologic data. The results indicate that the sedimentary sequence spans the interval from the Olduvai normal subchron to the end of the Brunhes normal chron. According to the lithology and the magnetic susceptibility record, the core can be divided into four intervals, from bottom to top: the Qian’an Formation (fluvial yellow-green, brown-red and grey-brown silty sand, interbedded with brown-black silty clay, with gravel layers in the lower part); the Lingzi Formation and Lindian Formation (lacustrine grey-green clay interbedded with grey-white silty sand, with fossil bivalves); and a loess-like soil (predominantly brown-yellow clayey silt, alternating with fluviolacustrine grey-white, brown and black clay). The magnetic susceptibilities (MS) of the sediments of the lower fluvial Qian’an Formation and upper loess-like soil strata are generally higher, indicating the neoformation of strongly magnetic minerals under predominantly oxidizing conditions. By contrast, lower MS values occur in the lacustrine sediments of the Lingzi Formation and Lindian Formation, which indicate a reducing environment. Overall, the MS record of the QAL core exhibits clear orbital-scale variations, with higher MS values in coarse-grained silt layers, and lower values in fine-grained clay layers. After the application of a 300 mT DC field, the isothermal remanence (IRM) is only 70%, and a hysteresis loop remained unclosed at 500 mT. These characteristics indicate the substantial presence of high-coercivity magnetic minerals in the intervals with low MS values. Thermomagnetic measurements ( χ - T ) curves show an abrupt decrease in the heating curve at 580°C, indicating that magnetite is the dominant magnetic mineral, and a further decrease above 580°C indicates the presence of hematite. After the application of an applied DC field of 300 mT, the IRM reached 90%, the hysteresis loop was closed, and the heating limb of the χ - T curve did not decrease significantly after 580°C, indicating that low-coercivity magnetic minerals contributed substantially to the magnetic properties within the intervals of higher MS. The lacustrine sediments of the core were dated to ~1180‒450 ka based on paleomagnetic and QSL dating, and interpolation of the age-depth relationship suggests that Songnen paleo-lake existed for ~730 ka. The grain-size record of the loess in the northeastern monsoon region shows no evidence of drought events at 450 ka, and records of MS and FeD/FeT from the deposits of the loess Plateau indicate a trend of increasing summer monsoon precipitation at this time. Therefore, the disappearance of the paleo-lake may be related to the multiple episodes of subsidence of the Yishu fault, which caused a diversion of the discharge of the Songhua River from the Songnen paleo-lake to the Sanjiang Plain. The high-resolution MS record and the precise chronology obtained in this study indicate that the lacustrine sedimentary sequence of QAL core can potentially resolve orbital-scale paleoclimate changes during the middle Pleistocene in mid-latitude East Asia. The Mid-Pleistocene Transition (MPT) is one of the most important climatic events in Quaternary paleoclimatology, and it is necessary to understand the response of different climatic systems to the MPT. Detailed environmental magnetic, sedimentological and geochemical studies of lacustrine sedimentary strata in the Northeast Plain in China should be carried out to explore the paleoclimatic record of the area, including the regional climatic response to the MPT.

Mid-Cretaceous Tuor-Yuryakh Section of Kotelnyi Island, New Siberian Islands: How Does the Probable Basement of Sedimentary Cover of the Laptev Sea Look on Land?

The model of geological structure of sedimentary cover of the Laptev Sea accepted by most geologists suggests that the lower seismic complex of the cover begins by the Aptian–Albian sedimentary rocks. They can be studied in natural outcrops of Kotelnyi Island. The section of the Tuor-Yuryakh Trough, which exposes the lower part of the Cretaceous complex, is described in the paper. It is composed of continental coaliferous rocks ~100 m thick. The marking beds divide it into five members, which are traced along the western wall of the trough at the distance up to 3 km. The spore–pollen complexes and plant megafossils indicate that almost the entire visible section of the mid-Cretaceous is Albian. Only its lower part no more than 14 m thick can probably belong to the Aptian. Marine facies with Albian foraminifers were found 15 m above the bottom of the Cretaceous complex. The section of the Cretaceous rocks is underlain by the Lower Jurassic marine clays and siltstones. The foraminifer assemblages of this part of the section are typical of the upper Sinemurian–Pliensbachian and fossil bivalves indicate late Sinemurian age of the host rocks. The hiatus ~70 Ma duration has no expression in the section and this boundary can de facto be substantiated only by microfossils. This vague contact between the Lower Jurassic and mid-Cretaceous rocks does not correspond to geophysical characteristics of the bottom of the lower seismic complex of the cover of the eastern part of the Laptev Sea. The latter is described as the most evident seismic horizon of the section of the cover, suggesting unconformable occurrence of the lower seismic complex on a peneplenized surface of lithified and dislocated rocks. This is mostly similar to the bottom of the Eocene sediments, which were observed on Belkovsky and Kotelnyi islands. The paper discusses possible application of our land results for interpretation of the shelf seismic sections of the Laptev Sea. It is concluded that local reasons are responsible for a vague boundary between the Lower Jurassic and mid-Cretaceous sequences in the section studied. Our observations support ideas on possible Aptian–Albian age of the rocks of the basement of the lower seismic complex; however, it is proposed to use also the previously popular idea on the Eocene age of the lower seismic complex of sedimentary cover of the eastern part of the Laptev Sea as one of the possible working scenarios.

Low drilling frequency in Norian benthic assemblages from the southern Italian Alps and the role of specialized durophages during the Late Triassic

Drillholes represent one of the clearest lines of evidence for predation of benthic invertebrates in the fossil record and are frequently used as a primary proxy for predation intensity in the fossil record. Drillholes are abundant in the late Cretaceous and Cenozoic, but their occurrence is patchy in older deposits of the Mesozoic. The inconsistent record of drillholes in pre-Cretaceous deposits of Mesozoic age are problematic for interpretations of predation-prey dynamics and adaptive radiations, and the role of taphonomy or diagenesis have not been resolved. Here we present drilling percentages for assemblages of well-preserved shelly benthic invertebrates (mainly comprised of bivalves and rare gastropods) from the upper Norian (Upper Triassic) in northern Italy in order to compare these values with reported drilling percentages from the Carnian San Cassiano Formation, a rare Triassic sedimentary unit that has yielded many drilled fossils. The Norian fossil deposits reported here are comparable to those of the San Cassiano in terms of depositional environment, preservation, and region, and can be reasonably compared to the drilling percentage of fossils from the San Cassiano. The sampled deposits are collected from marly limestone horizons in the Argillite di Riva di Solto in the Southern Italian Alps, deposited in the Lombardian Basin, and which are interbedded with shale units containing well-preserved fish and arthropod fossils, enabling a correlation between paleoecological structure of the shelly benthos and the demersal-pelagic predator diversity. Over four hundred bivalve fossils yielded a drilling percentage of 0.24% (1/406), which is typical for fossil assemblages of this age, but the single occurrence of a drillhole in this study is in marked contrast to the many drilled specimens reported from the San Cassiano Formation deposit in Italy. The drilled specimen (with complete drillhole) was an infaunal bivalve and no incomplete drillholes were observed in other specimens. Thus, drilling percentages for the Triassic are consistently low, but present, suggesting that drilling predation was an ecologically minimal influence to benthic communities and unlikely to have driven the significant ecological changes observed in benthic communities during the Late Triassic. Although drilling predation occurred during the Late Triassic, we present an updated database of specialized durophagous predators (including fishes, sharks, and reptiles) that are likely to have been more ecologically influential on benthic communities during the Norian Stage, fishes in particular.