Emergence Of New Levels Research Articles

Stability of ecologically scaffolded traits during evolutionary transitions in individuality

Evolutionary transitions in individuality are events in the history of life leading to the emergence of new levels of individuality. Recent studies have described an ecological scaffolding scenario of such transitions focused on the evolutionary consequences of an externally imposed renewing meta-population structure with limited dispersal. One difficulty for such a scenario has been explaining the stability of collective-level traits when scaffolding conditions no longer apply. Here, we show that the stability of scaffolded traits can rely on evolutionary hysteresis: even if the environment is reverted to an ancestral state, collectives do not return to ancestral phenotypes. We describe this phenomenon using a stochastic meta-population model and adaptive dynamics. Further, we show that ecological scaffolding may be limited to Goldilocks zones of the environment. We conjecture that Goldilocks zones—even if they might be rare—could act as initiators of evolutionary transitions and help to explain the near ubiquity of collective-level individuality.

Spontaneous Emergence of Multicellular Heritability.

The major transitions in evolution include events and processes that result in the emergence of new levels of biological individuality. For collectives to undergo Darwinian evolution, their traits must be heritable, but the emergence of higher-level heritability is poorly understood and has long been considered a stumbling block for nascent evolutionary transitions. Using analytical models, synthetic biology, and biologically-informed simulations, we explored the emergence of trait heritability during the evolution of multicellularity. Prior work on the evolution of multicellularity has asserted that substantial collective-level trait heritability either emerges only late in the transition or requires some evolutionary change subsequent to the formation of clonal multicellular groups. In a prior analytical model, we showed that collective-level heritability not only exists but is usually more heritable than the underlying cell-level trait upon which it is based, as soon as multicellular groups form. Here, we show that key assumptions and predictions of that model are borne out in a real engineered biological system, with important implications for the emergence of collective-level heritability.

Historical aspects of the development of technical diagnostics

The prerequisites for the formation and development of technical diagnostics as a science contain several stages, starting from the appearance of the first mechanisms, and then steam engines. The use of organoleptic methods for assessing the state (listening to noise, tactile analysis, visualization of mechanical vibrations, etc.), in addition to the developed safety devices and readings of instruments for measuring speed, pressure, temperature, ensured the functioning of mechanisms until the middle of the twentieth century. The emergence of complex technical systems associated with the implementation of atomic, space and other projects required ensuring their reliability with an insufficient level of data, which became the main reason for the emergence of technical diagnostics as a science. When determining the state of mechanical equipment, vibration has always been the main diagnostic parameter, but only with the development of microprocessor technology and computer technology has it become possible to obtain objective data. The complication of equipment and control systems leads to the emergence of new levels of obtaining and collecting diagnostic data, which is reflected in the appearance of large amounts of data and the need to form information flows for effective management of the technical condition of the equipment. Pattern recognition methods, after adapting to specific practical situations, allow you to get good results after laborious work by highly qualified specialists.

On the Theory of Narrative Levels and Their Annotation in the Digital Context

The article was written in the context of a Shared Task on the Analysis of Narrative levels Through Annotation (“SANTA”) which was published as a first draft in 2019. This revised version is based on further discussion on the formalization of the narratological concept of ‘narrative level.’ We firstly discuss the theory of narrative levels in literary studies, secondly derive features for the identification of narrative levels and finally develop guidelines for their annotation. An essential finding of the theoretical work lies in connecting the concept of ‘narrative level’ to the narrator. By identifying different types of narrators, we are able to enumerate and categorize different scenarios for the emergence of new levels in narrative texts. Hereby, the article does not remain restricted to prototypical cases, but also deals with rare and problematic cases. Overall, our goal is to provide a theoretical reflection on narrative levels and to create accurate guidelines for its recognition. The method of approaching the phenomenon through annotation has proven to be extremely fruitful particularly in identifying the boundaries of the narrative levels.

Scaffolding Natural Selection

Darwin provided us with a powerful theoretical framework to explain the evolution of living systems. Natural selection alone, however, has sometimes been seen as insufficient to explain the emergence of new levels of selection. The problem is one of “circularity” for evolutionary explanations: how to explain the origins of Darwinian properties without already invoking their presence at the level they emerge. That is, how does evolution by natural selection commence in the first place? Recent results in experimental evolution suggest a way forward: Paul Rainey and his collaborators have shown that Darwinian properties could be exogenously imposed via what they call “ecological scaffolding.” This could solve the “black box” dilemma faced by Darwinian explanations of new levels of organization. Yet, despite “scaffolding” recently becoming a popular term in the study of cognition, culture, and evolution, the concept has suffered from vagueness and ambiguity. This article aims to show that scaffolding can be turned into a proper scientific concept able to do explanatory work within the context of the major evolutionary transitions. Doing so will allow us to once again extend the scope of the Darwinian model of evolution by natural selection.

From Identity to Uniqueness: The Emergence of Increasingly Higher Levels of Hierarchy in the Process of the Matter Evolution.

This article focuses on several factors of complification, which worked during the evolution of our Universe. During the early stages of such evolution up to the Recombination Era, it was laws of quantum mechanics; during the Dark Ages it was gravitation; during the chemical evolution-diversification; and during the biological and human evolution—a process of distinctifying. The main event in the evolution of the Universe was the emergence of new levels of hierarchy, which together constitute the process of hierarchogenesis. This process contains 14 such events so far, and its dynamics is presented graphically by a very regular and smooth curve. The function that the curve presents is odd, i.e., symmetric about its central part, due to the similarity of patterns of the deceleration during the cosmic/chemical evolution (1st half of the general evolution) and the acceleration during the biological/human evolution (its 2nd half). The main driver of the hierarchogenesis as described by this odd function is counteraction and counterbalance of attraction and repulsion that take various forms at the different hierarchical levels. Direction and pace of the irreversible and inevitable increase of the Universe complexity in accordance with the general law of complification result from a consistent influence of all these factors.

Viruses and mobile elements as drivers of evolutionary transitions.

The history of life is punctuated by evolutionary transitions which engender emergence of new levels of biological organization that involves selection acting at increasingly complex ensembles of biological entities. Major evolutionary transitions include the origin of prokaryotic and then eukaryotic cells, multicellular organisms and eusocial animals. All or nearly all cellular life forms are hosts to diverse selfish genetic elements with various levels of autonomy including plasmids, transposons and viruses. I present evidence that, at least up to and including the origin of multicellularity, evolutionary transitions are driven by the coevolution of hosts with these genetic parasites along with sharing of ‘public goods’. Selfish elements drive evolutionary transitions at two distinct levels. First, mathematical modelling of evolutionary processes, such as evolution of primitive replicator populations or unicellular organisms, indicates that only increasing organizational complexity, e.g. emergence of multicellular aggregates, can prevent the collapse of the host–parasite system under the pressure of parasites. Second, comparative genomic analysis reveals numerous cases of recruitment of genes with essential functions in cellular life forms, including those that enable evolutionary transitions.This article is part of the themed issue ‘The major synthetic evolutionary transitions’.

Evolutionary biosemiotics and multilevel construction networks.

In contrast to the traditional relational semiotics, biosemiotics decisively deviates towards dynamical aspects of signs at the evolutionary and developmental time scales. The analysis of sign dynamics requires constructivism (in a broad sense) to explain how new components such as subagents, sensors, effectors, and interpretation networks are produced by developing and evolving organisms. Semiotic networks that include signs, tools, and subagents are multilevel, and this feature supports the plasticity, robustness, and evolvability of organisms. The origin of life is described here as the emergence of simple self-constructing semiotic networks that progressively increased the diversity of their components and relations. Primitive organisms have no capacity to classify and track objects; thus, we need to admit the existence of proto-signs that directly regulate activities of agents without being associated with objects. However, object recognition and handling became possible in eukaryotic species with the development of extensive rewritable epigenetic memory as well as sensorial and effector capacities. Semiotic networks are based on sequential and recursive construction, where each step produces components (i.e., agents, scaffolds, signs, and resources) that are needed for the following steps of construction. Construction is not limited to repair and reproduction of what already exists or is unambiguously encoded, it also includes production of new components and behaviors via learning and evolution. A special case is the emergence of new levels of organization known as metasystem transition. Multilevel semiotic networks reshape the phenotype of organisms by combining a mosaic of features developed via learning and evolution of cooperating and/or conflicting subagents.

Management of neck metastases in head and neck cancer: United Kingdom National Multidisciplinary Guidelines.

This is the official guideline endorsed by the specialty associations involved in the care of head and neck cancer patients in the UK. A rational plan to manage the neck is necessary for all head and neck primaries. With the emergence of new level 1 evidence across several domains of neck metastases, this guideline will identify the evidence-based recommendations for management. Recommendations • Computed tomographic or magnetic resonance imaging is mandatory for staging neck disease, with choice of modality dependant on imaging modality used for the primary site, local availability and expertise. (R) • Patients with a clinically N0 neck, with more than 15-20 per cent risk of occult nodal metastases, should be offered prophylactic treatment of the neck. (R) • The treatment choice of for the N0 and N+ neck should be guided by the treatment to the primary site. (G) • If observation is planned for the N0 neck, this should be supplemented by regular ultrasonograms to ensure early detection. (R) • All patients with T1 and T2 oral cavity cancer and N0 neck should receive prophylactic neck treatment. (R) • Selective neck dissection (SND) is as effective as modified radical neck dissection for controlling regional disease in N0 necks for all primary sites. (R) • SND alone is adequate treatment for pN1 neck disease without adverse histological features. (R) • Post-operative radiation for adverse histologic features following SND confers control rates comparable with more extensive procedures. (R) • Adjuvant radiation following surgery for patients with adverse histological features improves regional control rates. (R) • Post-operative chemoradiation improves regional control in patients with extracapsular spread and/or microscopically involved surgical margins. (R) • Following chemoradiation therapy, complete responders who do not show evidence of active disease on co-registered positron emission tomography-computed tomography (PET-CT) scans performed at 10-12 weeks, do not need salvage neck dissection. (R) • Salvage surgery should be considered for those with incomplete or equivocal response of nodal disease on PET-CT. (R).

Synthesis of carbon doped WO3·0.33H2O hierarchical photocatalyst with improved photocatalytic activity

A carbon doping method is developed to overcome the drawback of conventional carbon doping method. The carbon dopant is synthesized by treating carbon with dilute nitric acid through hydrothermal process. Carbon-doped WO3·0.33H2O (CWO3·0.33H2O) is obtained by adding the carbon dopant solution, and then through an additional hydrothermal process. Based on XRD, Raman and XPS, carbon have been incorporated into the interstitial position of WO3·0.33H2O unit cell rather than substitution of oxygen in lattice of WO3·0.33H2O, which induces distortion of WO6 octahedron and lattice defect. In addition, carbon doping also induces the emergence of new level above the valence band. The new level increases the migration pathway of photo-generated carriers, which reduces the recombination rate of photo-generated carriers. The changes of band structure, distortion of WO6 octahedron and lattice defect are beneficial to improve the photocatalytic performance of WO3·0.33H2O. Compared with pristine WO3·0.33H2O, CWO3·0.33H2O display better photocatalytic performance on degradation of rhodamine B (RhB), which include the faster de-ethylation process and cycloreversion process. Moreover, the experiments of radical quenchers confirmed that the h+ display the main influence on degradation of RhB. This study implies that carbon can be doped into some metal oxide hydrates through a mild preparation process, which provides a possible way for the synthesis of carbon doped thermal unstable materials.

Human Metasystem Transition (HMST) Theory

Metasystem transitions are events representing the evolutionary emergence of a higher level of organization through the integration of subsystems into a higher “metasystem” (A1+A2+A3=B). Such events have occurred several times throughout the history of life (e.g., emergence of life, multicellular life, sexual reproduction). The emergence of new levels of organization has occurred within the human system three times, and has resulted in three broadly defined levels of higher control, producing three broadly defined levels of group selection (e.g., band/tribe, chiefdom/kingdom, nation-state/international). These are “Human Metasystem Transitions” (HMST). Throughout these HMST several common system-level patterns have manifested that are fundamental to understanding the nature and evolution of the human system, as well as our potential future development. First, HMST have been built around the control of three mostly distinct primary energy sources (e.g., hunting, agriculture, industry). Second, the control of new energy sources has always been achieved and stabilized by utilizing the evolutionary emergence of a more powerful information-processing medium (e.g., language, writing, printing press). Third, new controls emerge with the capability of organizing energy flows over larger expanses of space in shorter durations of time: bands/tribes controlled regional space and stabilized for hundreds of thousand of years, chiefdoms/kingdoms controlled semi-continental expanses of space and stabilized for thousands of years, and nation-states control continental expanses of space and have stabilized for centuries. This space-time component of hierarchical metasystem emergence can be conceptualized as the active compression of space-time-energy-matter (STEM compression) enabled by higher informational and energetic properties within the human system, which allow for more complex organization (i.e., higher subsystem integration). In this framework, increased information-energy control and feedback, and the consequent metasystem compression of space-time, represent the theoretical pillars of HMST theory. Most importantly, HMST theory may have practical application in modeling the future of the human system and the nature of the next human metasystem.

Development (and Evolution) of the Universe

I distinguish Nature from the World. I also distinguish development from evolution. Development is progressive change and can be modeled as part of Nature, using a specification hierarchy. I have proposed a ‘canonical developmental trajectory’ of dissipative structures with the stages defined thermodynamically and informationally. I consider some thermodynamic aspects of the Big Bang, leading to a proposal for reviving final cause. This model imposes a ‘hylozooic’ kind of interpretation upon Nature, as all emergent features at higher levels would have been vaguely and episodically present primitively in the lower integrative levels, and were stabilized materially with the developmental emergence of new levels. The specification hierarchy’s form is that of a tree, with its trunk in its lowest level, and so this hierarchy is appropriate for modeling an expanding system like the Universe. It is consistent with this model of differentiation during Big Bang development to view emerging branch tips as having been entrained by multiple finalities because of the top-down integration of the various levels of organization by the higher levels.

The cyborg, the ideology chip and the guru programme: the implications of cyborg technologies for the development of human consciousness

There has been little thought given in science to the impact of direct brain‐machine interfacing upon the future development of human consciousness. Even less thought has been given to the possibilities for both optimizing and thwarting development in the cyborg child. A neurocognitive model of the evolution of cyborg consciousness is summarized, and from this model grounded speculations are offered pertaining to the future development of the higher cognitive functions in the cyborg child. It will be shown that cybernetic implants are “multistable”; that is, the artificial intelligence (AI) component of the cyborg brain‐machine linkage may function to condition development along ideological lines (the brain conditioned by the “ideological chip”), or may operate to open up neurocognitive development to new and heretofore unrealized limits (the brain’s development optimized by the “guru programme”). Development of the cyborg child may be conditioned in the interests of ideological concerns, or may lead to a consciousness that easily transcends all forms of ideology. Application of the guru programme may foster the emergence of new levels of cognitive complexity and information processing (à la Piagetian and neo‐Piagetian theory) that in turn allows new strategies of adaptation previously beyond human comprehension. The ethical and regulatory problems raised by cyborg technologies are addressed.

Opening up closure. Semiotics across scales

The dynamic emergence of new levels of organization in complex systems is related to the semiotic reorganization of discrete/continuous variety at the level below as continuous/discrete meaning for the level above. In this view both the semiotic and the dynamic closure of system levels is reopened to allow the development and evolution of greater complexity.

Equilibrium configurations of N equal charges on a sphere

The emergence of new levels of complexity that often accompanies the transition from few- to many-body systems is clearly illustrated by the progression of equilibrium states of N charges on the surface of a sphere as N increases. The characteristics of these electrostatic equilibrium states in the range 2<or=N<or=65 can be examined in detail by equilibrating 1000 randomized initial configurations for every value of N. As N increases, the equilibrium states undergo a succession of structural changes. For instance, a state with non-zero dipole moment appears at N=11, an enantiomeric or mirror image state appears at N=15, and a robust metastable state appears at N=16. For values of N exceeding 50, clusters of four or more metastable states with energies within 0.01% of each other are the dominant pattern. In analogy with some other complex systems, these energetically similar states have strikingly different asymmetric configurations.

Emergent attractors and the law of maximum entropy production: Foundations to a theory of general evolution

AbstractBy Boltzmann's widely accepted reduction of the second law to a stochastic collision function, transformations from an incoherent to a coherent state are ‘infinitely improbable’. Yet it is precisely the transformation of the incoherent into the coherent, the progressive emergence of new levels of macroscopic constraints, that has brought, and continues to bring, the visible universe into being. Whether in ecosystem succession, the emergence of hydrodynamic structures, morphogenesis, 4GY of bioevolution, cognitive and cultural evolution, or cosmic evolution as a whole of which all the rest are part, the observed behavior is the same: Spontaneous and successive symmetry‐breaking coupled with the progressive attraction of matter away from equilibrium to time‐independent limit sets through the selection of some highly reduced (coherent) subset of accessible microstates, Ms, from some much larger initial (incoherent) set, Mn. This immediately raises the question of where, given the known laws of physics, such progressive goal‐directed behavior can come from, viz., (i) what universality governs the instability (symmetry‐breaking) of the incoherent regime; and (ii) what extremum principle governs the selection of microstates on the attractor? It is shown that a law of maximum entropy production can be reached deductively from the second law of thermodynamics with an assumption of the ontological irreducibility of nonlinear behavior, and inductively from the study of simple physical systems where the usual explanatory mechanisms, e.g., genes, brains or other ‘maker’, are clearly absent and therefore cannot be invoked. This law is seen as the cornerstone to a theory of general evolution within which biological and cultural evolution are special cases. As a result, a great proportion of the behavior generally believed to be generic to either biological or cultural systems is shown instead to be generic to the physics of the expanding universe.