Effects Of Fire Frequency Research Articles

A probabilistic selection of fire scenarios considering operating condition in aircraft carrier’s hangar spaces

ABSTRACT Aircraft carrier hangars require sufficient space to enable essential functions such as aircraft preparation, stowage, handling of aviation fuel and munitions, etc. Consequently, the hangars are treated as high-fire-risk spaces owing to their operating conditions. Although hangar fire accidents are caused by various temporally and spatially varying fire hazards, fire safety designs for hangars follow a deterministic approach based on IMO regulations, similar to ro-ro ships. Hence, this paper proposes one hundred credible fire scenarios to enhance the fire safety design of hangar structures by utilizing Latin hypercube sampling. These fire scenarios were also employed to determine fire-induced loads in the form of exceedance curves. Moreover, the effect of fire frequency defining method and ship motions on fire consequences was quantitatively investigated through case study and CFD simulation.

The large‐scale restoration of fire and water regimes in Everglades National Park reveal little change in plant diversity along an elevational gradient

Quantifying the response of plant diversity to large‐scale restoration is essential for measuring management success. One of the world's largest restoration efforts began in 2000 in the Everglades ecosystem in Florida, United States, through the Comprehensive Everglades Restoration Plan. In coordination with ongoing fire management, this restoration effort aims to restore natural hydrologic and fire regimes in this dynamic ecosystem. Restored water and fire regimes in the Everglades interact along an elevational gradient between seasonally inundated marl prairie and frequently burned pine rockland on Long Pine Key in Everglades National Park. To determine the impact of management on plant community richness and composition, we resampled transects spanning the elevational gradient between marl prairie and pine rockland initially sampled prior to the implementation of current restoration efforts (circa 1997–1999). We measured percent plant cover and developed generalized linear mixed models to determine the effects of fire frequency, average water depth, and elevation on plant species richness and composition across the two time periods. Additionally, we used species‐level random effects to examine how individual species respond to each environmental variable. We failed to detect any systematic shifts in plant composition in response to fire and water management. However, we found that complex interactions between fire and water structure plant composition and maximize species richness along gradients of water depth and elevation, with low, moderate, and high burn frequencies increasing richness in wet, moderately wet, and dry sites, respectively.

Resilience to summer bushfire in the threatened orchid, Caladenia tessellata, in terms of pollination success, herbivory, and mycorrhizal associations

Abstract In some biogeographic regions, many threatened plant species occur in habitats that periodically experience bushfire. However, we currently have relatively little information on how important plant–animal and plant–fungus interactions are affected by these fires. For the threatened sexually deceptive orchid Caladenia tessellata we test whether pollination rate, frequency of florivory, and the species of mycorrhizae the plant associates with differ between burnt and unburnt sites. Interestingly, pollination rates were unaffected by fire, demonstrating that populations of the thynnine wasp pollinator can persist post-fire. However, there was a significant negative relationship between number of flowers in a population and pollination success, which is likely a by-product of a deceptive pollination strategy. Despite the presence of vertebrate herbivores, florivory rates were low in both burnt and unburnt sites. Caladenia tessellata associated primarily with the mycorrhiza Serendiptia australiana, regardless of fire history. While our results suggest resilience to a one-off summer fire for the ecological interactions that we measured, it would be interesting to investigate the effects of fire frequency and time of year. High reproductive rates in small populations of C. tessellata suggest these populations may be viable and that retaining them is a high priority for conservation.

Fire Has a Positive Effect on the Abundance of Sun Spiders (Arachnida: Solifugae) in the Cerrado-Pantanal Ecotone

Fire is an important disturbance factor in shrublands, grasslands, and savannas. It alters the habitat of a multitude of species and, under natural dynamics, is a major determinant of landscape vegetation patterns. Here, we evaluate the effects of different wildfire regimes on the abundance of sun spiders in the Cerrado-Pantanal ecotone. To study how different fire regimes affect the number of individual sun spiders, we considered the frequency of fire occurrences in the last 20 years and classified locations as high frequency or low frequency. We also classified the time of the last fire in 2020 as occurring in the first or second half of the year. In addition, we compared the number of individual sun spiders before and after fire. We found no effects of fire frequency and period when the fire occurred in 2020, but the number of individual sun spiders was higher after wildfires. Although ground-dwelling are considered fire sensitive, some can employ strategies to tolerate fire so that they are able to not only survive, but also reproduce in fire-prone landscapes. Thus, we suggest that sun spiders are resilient, can explore sites under different fire regimes, and can be considered pyrophilous species.

Should we burn the Cerrado? Effects of fire frequency on open savanna plant communities

AbstractAimsTropical savanna evolved in the presence of fire, depending on it to maintain its physiognomies and biodiversity. Open ecosystems of tropical savanna burn frequently, and thus changes in fire frequency can affect their plant communities. In this study we aimed to analyze the effects of different fire frequencies on the composition, structure and post‐fire regeneration of open savanna plant communities of the Cerrado.LocationOpen savannas, Central Brazil (13°35–38′S and 47°45–51′W).MethodsWe conducted surveys in areas with different fire frequencies (fire exclusion, burned annually and burned biennially for 6 years, from 2013). Vegetation sampling was carried out before application of fire treatments in 2017, and then 1 and 2 years after. Species cover was estimated visually and later grouped into graminoids, shrubs and forbs.ResultsEvaluation of the plant communities after 6 years of fire treatment indicated that fire frequency did not affect species richness. Fire frequency did, however, promote alteration in species occurrence and abundance, resulting in three dissimilar plant communities. Regarding post‐fire regeneration, annually burned plots showed consistency in species richness and growth form cover over 6 years of annually applied fire. Biennially burned plots tended to be more variable than the other treatments, with a large increase in species richness and growth form cover in the first year after fire, succeeded by a decrease in the second year. Not all species that were recruited just after fire were able to persist in the above‐ground community in the second year after fire.ConclusionsOpen savannas of the Cerrado are highly resilient to fire, even to annual fires, with 6 years of annual fire treatments not affecting post‐fire species diversity or growth form cover. However, communities differed in species composition, which can influence their functional diversity and lead to alterations in ecosystem services. Our findings encourage long‐term studies to evaluate the resilience of areas frequently burned, as well as longer fire exclusion periods.

Extending the National Burned Area Composite Time Series of Wildfires in Canada

Wildfires are a major natural disturbance in Canada that are postulated to increase under a warming climate. To derive accurate trends in burned area and to quantify the effects of fire frequency, duration, and extent, a sufficiently long time series of reliable burned area maps is required. With that in mind, we extended Canada’s National Burned Area Composite (NBAC) dataset from its previous start year (2004) back to 1986. NBAC consists of annual maps in polygon format where the area burned in each fire event is represented by the best available delineation among various mapping methods and data sources of varying quality. Ordered from more to less reliability, in the new 35-year time series (1986 to 2020), 10% of the total burned area was derived from airborne and high-resolution (<5 m) satellite imagery, 81% from change detection methods using 30 m Landsat satellite imagery, and the remaining 9% was largely from aerial surveys. Total (Canada-wide) annual burned area estimates ranged from 215,797 ha in 2020 to 6.7 million ha (Mha) in 1989. We computed 95% confidence intervals for the estimate of each year from 1986 to 2020 based on the accuracy and relative contribution in that year of the different data sources, for both the new NBAC time series and the polygon version of the Canadian National Fire Database (CNFDB), a commonly used source of spatially explicit data on burned area in Canada. NBAC confidence intervals were on average ±9.7% of the annual figure, about one-third the width of the confidence intervals derived for CNFDB. The NBAC time series also included nearly 5000 fire events (totalling 4 Mha, with the largest event being 120,661 ha in size) that are missing in the CNFDB. In a regional analysis for the Northwest Territories, retroactive fire mapping from Landsat imagery reduced historical estimates by 3 Mha (16%), which would result in a 1.6 Mha increase in the reported undisturbed critical habitat for threatened woodland caribou. The NBAC dataset is freely downloadable from the Canadian Wildland Fire Information System.

Remarkable Resilience of Forest Structure and Biodiversity Following Fire in the Peri-Urban Bushland of Sydney, Australia

In rapidly urbanizing areas, natural vegetation becomes fragmented, making conservation planning challenging, particularly as climate change accelerates fire risk. We studied urban forest fragments in two threatened eucalypt-dominated (scribbly gum woodland, SGW, and ironbark forest, IF) communities across ~2000 ha near Sydney, Australia, to evaluate effects of fire frequency (0–4 in last 25 years) and time since fire (0.5 to >25 years) on canopy structure, habitat quality and biodiversity (e.g., species richness). Airborne lidar was used to assess canopy height and density, and ground-based surveys of 148 (400 m2) plots measured leaf area index (LAI), plant species composition and habitat metrics such as litter cover and hollow-bearing trees. LAI, canopy density, litter, and microbiotic soil crust increased with time since fire in both communities, while tree and mistletoe cover increased in IF. Unexpectedly, plant species richness increased with fire frequency, owing to increased shrub richness which offset decreased tree richness in both communities. These findings indicate biodiversity and canopy structure are generally resilient to a range of times since fire and fire frequencies across this study area. Nevertheless, reduced arboreal habitat quality and subtle shifts in community composition of resprouters and obligate seeders signal early concern for a scenario of increasing fire frequency under climate change. Ongoing assessment of fire responses is needed to ensure that biodiversity, canopy structure and ecosystem function are maintained in the remaining fragments of urban forests under future climate change which will likely drive hotter and more frequent fires.

The effects of fire frequency on vegetation structure and mammal assemblages in a savannah‐woodland system

AbstractFire frequency affects vegetation structure and composition in savannah‐ woodlands, with the potential indirect consequence of affecting the habitat choice and space use of large mammals. Understanding these responses can assist with the development of appropriate fire management policies to conserve co‐existing species with different habitat requirements. In a natural experiment in Majete Wildlife Reserve, Malawi, we compared vegetation composition, woody plant structure and large (>5 kg) mammal assemblages on 10 frequently and 10 infrequently burnt 30‐ha plots, that had fire return intervals of <2.5 and >6 years, respectively, between 2001 and 2019. Fire frequency had little effect on woody plant composition, but did affect grass species composition. Frequent burning reduced mean tree height from 4 to 3 m, woody plant cover from 59% to 35%, and resulted in the dominance of short‐statured, less‐palatable grasses. Many mammal species showed clear preferences for either frequently burnt (e.g. buffalo, zebra, kudu, impala and sable) or infrequently burnt (e.g. nyala, bush pig, baboon, common duiker and vervet monkey) areas, while others (e.g. elephants and black rhinoceros) were not selective. We conclude that establishing a mosaic of patches exposed to different fire frequencies would allow for the persistence of all mammal species in the area by creating an optimal range of habitat types.

Anthropogenic fire practices only ‘best’ if they promote a large seed buildup: comment on the conservation needs of a fire-killed grevillea

Fire regimes control the population dynamics of fireprone plant species. Here, I re-analyse the data acquired in a study of the conservation requirements of a rare, fire-killed grevillea. I conclude that specific knowledge is required on the rate at which seeds accumulate in the soil, and the effect of fire frequency on population dynamics for a comprehensive understanding of the conservation biology of this species. Modelling indicates that fires at intervals in the order of multiple decades and covering at least 60% of the ground surface are optimal for this species, independently of the type of fire employed.

How and why grasshopper community maturation rates are slowing on a North American tall grass prairie.

Invertebrate growth rates have been changing in the Anthropocene. We examine rates of seasonal maturation in a grasshopper community that has been declining annually greater than 2% a year over 34 years. As this grassland has experienced a 1°C increase in temperature, higher plant biomass and lower nutrient densities, the community is maturing more slowly. Community maturation had a nutritional component: declining in years/watersheds with lower plant nitrogen. The effects of fire frequency were consistent with effects of plant nitrogen. Principal components analysis also suggests associated changes in species composition-declines in the densities of grass feeders were associated with declines in community maturation rates. We conclude that slowed maturation rates-a trend counteracted by frequent burning-likely contribute to long-term decline of this dominant herbivore.

Long-Term Effect of Fire Frequency on Miombo Woody Vegetation Dynamics: The Case of Niassa Special Reserve, Northern Mozambique

Long-Term Effect of Fire Frequency on Miombo Woody Vegetation Dynamics: The Case of Niassa Special Reserve, Northern Mozambique

Effect of Fire Frequency on the Flammability of Two Mediterranean Pines: Link with Needle Terpene Content

Flammability is a major factor involved in Mediterranean plant evolution that has led to the diversity of fire-related traits according to fire regimes and fire-adaptive strategies. With on-going climate change, new fire regimes are threatening plant species if they do not adapt or acclimate. Studying flammability and terpene content variation according to the different fire frequencies in the recent fire history represents a great challenge to anticipating the flammability of ecosystems in the near future. The flammability of shoots and litter as well as the needle terpene contents of two pine species with different fire adaptive strategies (Pinus halepensis and Pinus sylvestris) were measured according to two fire modalities (0 vs. 1–2 fire events over the last 60 years). Results showed that, regardless of the species and the fuel type, flammability was higher in populations having undergone at least one past fire event even when factors influencing flammability (e.g., structural traits and hydric content) were considered. The terpene content did not vary in P. sylvestris’ needles according to the fire modality, but that of sesqui- and diterpenes was higher in P. halepensis’ needles sampled in the “Fire” modality. In addition, associations made between flammability and terpene content using random forest analyses indicated that the terpene molecules differed between fire modalities for both species and fuel types. The same results were obtained with significant terpenes driving flammability as were highlighted in the PLS analyses, especially for P. halepensis for which enhanced shoot flammability in the “Fire” modality agreed with the adaptive strategy of this species to fire.

Influence of burning and defoliation on Festuca costata (Nees) in the Drakensberg

Fire occurs naturally in grasslands and savannas and it is commonly used as a management tool to influence structure and composition. The ability of a grass plant to recover from fire may be influenced by the fire frequency and intensity (influenced by seasonal weather conditions and fuel load), defoliation and competition from neighbouring plants. This study sought to investigate factors influencing the fire and herbivore dynamics of an evergreen C3 grass, Festuca costata, in a high-altitude C4 dominated grassland in the Drakensberg Mountains of South Africa. We experimentally examined the effect of fire frequency, defoliation and competition from neighbours on the growth dynamics of F. costata. Annual burning significantly decreased aboveground biomass production, number of tillers and tuft circumference of F. costata. Defoliation similarly reduced aboveground biomass production, number of tillers, tuft circumference and specific leaf area of F. costata. Competition from neighbouring plants reduced aboveground biomass production, number of tillers and tuft circumference of F. costata. Given the expected variable impacts of increasing CO2 levels and temperature on C3 and C4 grasses, these results provide insights for managing the matrix of C3 and C4 grasses in the Drakensberg grasslands in a changing climate.

Recovery of Chilean Mediterranean vegetation after different frequencies of fires

Post-fire recovery of sites in the Chilean Mediterranean vegetation were evaluated 20–30 years after the last fire. We mapped all fires that occurred between 1985 and 2015 in Central Chile using Landsat images. In order to conduct a spatial analysis of vegetation recovery and field sample, we chose sites burned between 1985 and 1995 that retained native vegetation and cover until 2015. In a sampled of these sites, richness and abundance of woody vegetation, and herbaceous richness were recorded. We contrasted our results from field sampling with control (unburned) sites at the species level. Generalized linear models (GLM) were used to evaluate the relationship between the percentage of vegetation recovered with fire frequency, pre-fire cover, topographic and geographic factors. In addition, GLM were used to evaluate the effects of fire frequency on species richness, abundance, and cover. We found that the proportion of dense and semi-dense vegetation cover were similar in sites burned once and twice, and higher than sites burned three times. Besides, the proportion of dense and semi-dense vegetation cover were higher in lower elevation sites, in those with higher slopes, and far from population centers. The richness and abundance of adult woody species, richness of regeneration and richness of native herbs, were greater in sites that had lesser fire frequency. Mean species richness of native herbaceous species decreased as fire frequency increased, exotic herbaceous cover had no relation with fire frequency, and both native and exotic herbaceous cover were greater at driest latitudes but were not related to fire frequency. Mean species richness after one fire was higher than in unburned sites, but some plant species were found only at unburned sites. We conclude that the Mediterranean vegetation of Chile is able to regenerate to pre-fire conditions after one and two fires, but three consecutive fires reduced its cover, richness and abundance, even 20 years after the fire.

Fire frequency effects on cleistogamy expression and progeny performance in Cologania broussonetii.

Increased fire frequency usually erodes microenvironmental conditions, causing a drastic limitation of edaphic resources. Thus, the production of permanently closed-small flowers (cleistogamous, CL) should increase in sites with high fire frequency as this implies a less expensive reproductive assurance strategy. However, because open, insect-pollinated flowers (chasmogamous, CH) have the potential capacity to outcross via pollinators, CH progeny produced at any site should outperform selfed CL progeny. We evaluate the effect of fire frequency on the relative production of CL/CH flowers and fruits, and their seed set, along with several progeny performance parameters in Cologania broussonetii (Fabaceae), a resprouting herb with dimorphic cleistogamy native to the Chaco Serrano. Fire frequency increased cleistogamy expression, reaching extreme levels in high fire frequency sites. Seed set was similarly high for both CH and CL flowers in the unburned condition, while in burned sites the few developed CH flowers set more seeds than CL flowers. However, progeny performance was similar between CH and CL progeny at each and across all fire frequency conditions. Cleistogamy expression in C. broussonetii is maximized in abiotically degraded frequently burned habitats, although the selfed CL progeny is as successful as potentially outcrossed CH progeny. Fire frequency may decreased floral size and abundance, selecting for autogamous reproduction, which restricts not only the genetic potential of plant populations but also the resources offered to pollinators. At the community level, increased cleistogamy expression may potentially have negative implications for non-cleistogamous, more outcrossing species surviving in frequently burned environments.

Effects of Fire Frequency on Woody Plant Composition and Functional Traits in a Wet Savanna Ecosystem

The objective of this study was to assess the effect of fire frequency on vegetation taxonomic and functional diversity in a wet savanna ecosystem, eastern Zimbabwe. The study area was stratified into three fire frequency regimes using a 15-year fire history (2000–2014) across the landscape: high (HFF: burnt every 1-2 years), medium (MFF: burnt every 3-4 years), and low (LFF: burnt every 5-6 years). Data were collected from a total of 30 plots measuring 20 m × 20 m each between March and May 2018. In each plot, we recorded tree maximum height (Hmax), woody plant density, basal diameter, resprouting capacity, and bark thickness. We calculated species evenness, diversity, functional richness (FRic), Rao’s Quadratic Entropy (RaoQ), functional redundancy, and relative bark thickness. We recorded 1,031 individual trees belonging to 24 species across the three fire regimes. Significant differences across the three fire regimes were recorded for Hmax, woody plant density, and relative bark thickness P < 0.05 . Hmax and woody plant density were higher in LFF than HFF regimes while relative bark thickness was higher in HFF than in the LFF regimes. Species evenness was significantly higher in HFF and MFF regimes than LFF regime P < 0.05 , while FRic and functional redundancy significantly increased with decreasing fire frequency P < 0.05 . However, no significant differences were recorded for resprouting capacity, species richness, taxonomic diversity, and RaoQ P > 0.05 . Species like Cassia petersiana, Cussonia spicata, Vachellia spp., and Rhus lancea were associated with LFF, while species like Protea gaguedi, Brachystegia utilis, and Vangueria infausta showed a strong association with HFF to MFF. Our study demonstrated that a combination of taxonomic and functional diversity metrics is adequate to evaluate the response of savanna vegetation to fire. We recommend a further assessment on vegetation composition using other elements of fire regimes.

Effects of fire frequency on savanna butterfly diversity and composition: A preliminary study

Fire plays a major role in many biomes, is widely used as a management tool and is likely to be affected by climate change. For effective conservation management, it is essential to understand how fire regimes affect different taxa, yet responses of invertebrates are particularly poorly documented. We tested how different fire frequencies influence savanna butterfly diversity and composition by using a long-term savanna fire experiment initiated in 1954 in the Kruger National Park (South Africa). We compared butterfly abundance, species richness and community composition across three fire frequencies: high (burnt annually), medium (burnt triennially) and low (burnt twice in 60 years). Plots with high fire frequency hosted higher abundance than medium- or low-frequency plots. Fire frequencies did not affect species richness, but they led to distinct communities of butterflies. Our findings suggest that, in view of the three fire frequencies tested, a spatial diversity of fire frequencies may increase butterfly diversity at the landscape level in wet savannas. Managers may need to promote a greater diversity of fire frequencies by increasing fire frequency in some areas to provide habitat for species requiring high fire frequency, and by decreasing fire frequency in a large proportion of the landscape to provide fire refuges. This study provides new insights for butterfly conservation in savannas and highlights several knowledge gaps, which further studies should address for insect responses to be given adequate consideration in fire management strategies.Conservation implications: A spatial diversity of fire frequencies may increase butterfly diversity. Managers may need to promote a greater diversity of fire frequencies by increasing fire frequency in some areas to provide habitat for species requiring high fire frequency, and by decreasing fire frequency in other areas to provide fire refuges.

EFFECTS OF FIRE ON STRUCTURE AND COMPOSITION OF VEGETATION COMMUNITY IN PINE FORESTS, NAM NAO NATIONAL PARK, THAILAND

Anthropogenic forest fires, including those occurring in the pine forests of Thailand, have been widely reported. Although pine forests are fire dependent, too frequent burning could be unhealthy for the forest as it might result in nutrient depletion and ecosystem degradation. Hence, this study aimed to examine the effects of forest fire on the vegetation structure and composition of a degraded pine forest (PF) and in a mixed pine-oak forest (O-PF) of Phu Kum Khao, Nam Nao National Park, Phetchabun Province, Thailand. The effects of fire in one year were studied using three 50 x 50 m experimental plots established in each forest type. Results showed that fire events caused a significantly higher tree mortality rate in the degraded pine forest than in the mixed pine-oak forest. Moreover, the seedling ratio (number of sprout to number of seedling ratio) in the mixed pine-oak forest was higher than in the degraded pine forest. Although the sapling and seedling diameters were slightly different between the sites, no significant effect of burning was observed on the sapling and seedling density, diameter and height. This might be due to the high variation of fire behavior in each plot. Since forest fires are a potential factor that might cause forest degradation in the study area, further studies on the effects of fire frequency on vegetation structure and composition is needed to provide sufficient fire control measures for efficient forest fire management to prevent further ecosystem degradation.

Effects of Fire Frequency on Seed Sources and Regeneration in Southeastern Amazonia

The trajectory and recovery time of fire-disturbed forests depend on the capacity of seedlings and resprouts to get established over time. Here, we investigated the mechanisms associated with fire effects on postfire regeneration in the context of a large-scale fire experiment located in southeastern Amazonia. Specifically, we tested the hypothesis that recurrent understory fires reduce forest regeneration capacity by reducing the recruitment of seedlings, the viability of the soil seed bank, the seed rain, and the resprouts. The experiment was comprised of three 50-ha plots: an unburned control, and two plots that were experimentally burned three (triennially) or six (annually) times between 2004 and 2010. Seedlings represented 2.4% and 0.6% of the seed rain and soil seed bank in the Control and burned treatment plots, respectively. Compared to the control, the triennial burns caused more reduction than the annual ones in seed rain (-42% versus -10%) and seed bank (-78% versus -50%). These fire-related reduction in seed rain in both treatments explain most of the low postfire density of seedlings (0.4 and 0.8 ind/m2, respectively), compared with the Control (2,9 ind/m2). The three-year interval between fires contributed to highest mortality of seeds stored in the soil than in the annual burn treatment, where fires were less severe. However, there were 5.5 times more resprouting in the plot burned triennially compared with the one burned annually. In both treatments, though, the number of resprouts declined over time. In conclusion, seeds from both seed rain and seed bank and resprouting were negatively affected by both fire frequencies, reducing the likelihood of native species recovery by altering the regeneration pathways. This situation is expected to be increasingly common in southeastern Amazon forests, particularly under warmer and dryer climate conditions.

Fire frequency effects in a grassy woodland: Trees and grasses

AbstractOur study investigated the effects of fire frequency on the tree and grass components of Cumberland Plain Woodland, an endangered grassy eucalypt woodland in eastern Australia. We located three sites within each of three fire frequency classes (high, moderate, low) based on fire history and a similar time since last fire. For trees, we asked whether fire frequency affected density, eucalypt population structure, basal area and spatial patterning. For grasses, we tested for fire frequency effects on total abundance and the abundance of two common species. Density of trees was not significantly affected by fire frequency for juveniles, saplings or adult trees and neither was basal area per hectare. Features of a recruitment bottleneck model were present for eucalypts. There was a large pool of suppressed juveniles making up more than half the population at each fire frequency. Saplings were the smallest group and were susceptible to fire‐induced stem mortality, with particularly low numbers relative to juveniles and small trees where fire was frequent or intense. Despite this difference in sapling mortality, numbers of small trees did not differ with fire frequency. Saplings could transition to small trees in equal numbers if they did so more quickly at high than at lower fire frequencies, and if recruitment of saplings into the tree layer was controlled independently of fire frequency. The size hierarchy of small to medium eucalypt trees was homogeneous over eight of the nine sites, spatial patterning of adult trees was random tending to regular, and mean tree size decreased with density at all sites. These features of eucalypt population structure are indicative of possible resource competition which could regulate tree recruitment. Total grass cover index was high across all fire frequencies, with Themeda triandra dominating at high and moderate fire frequencies and Microlaena stipoides at low fire frequency.