Eastern Tropical Pacific Research Articles

Larval Transport Pathways for Lutjanus peru and Lutjanus argentiventris in the Northwestern Mexico and Tropical Eastern Pacific

Understanding how ocean currents influence larval dispersal and measuring its magnitude is critical for conservation and sustainable exploitation, especially in the Tropical Eastern Pacific (TEP), where the larval transport of rocky reef fish remains untested. For this reason, a lagrangian simulation model was implemented to estimate larval transport pathways in Northwestern Mexico and TEP. Particle trajectories were simulated with data from the Hybrid Ocean Coordinate Model, focusing on three simulation scenarios: (1) using the occurrence records of Lutjanus peru and L. argentiventris as release sites; (2) considering a continuous distribution along the study area, and (3) taking the reproduction seasonality into account in both species. It was found that the continuous distribution scenario largely explained the genetic structure previously found in both species (genetic brakes between central and southern Mexico and Central America), confirming that the ocean currents play a significant role as predictors of genetic differentiation and gene flow in Northwestern Mexico and the TEP. Due to the oceanography of the area, the southern localities supply larvae from the northern localities; therefore, disturbances in any southern localities could affect the surrounding areas and have impacts that spread beyond their political boundaries.

Reproductive biology of the rock oyster Striostrea prismatica in two populations of the Mexican Central Pacific

ABSTRACT The oyster Striostrea prismatica is a fishery-relevant species in the eastern tropical Pacific, particularly in the Mexican Central Pacific (MCP), and little is known about its reproductive cycle. Therefore, our study analysed gametogenesis, sex ratio, sexual maturity stages, shell height at first maturity, and condition index during 2014 and 2015 in El Tizate and La Victoria in the MCP. The sex ratio was significantly biased towards males at both sites, presumably due to ontogeny or fishing pressure. The highest condition index was observed between April and May in both years, while lower values were observed from September to December. The reproductive cycle in both sites was annual, with maturity and spawning from June to August and the degradation phase in August. The condition index showed a negative correlation with temperature and chlorophyll a in both sites: as a result, the period of maximum maturity and spawning coincided with the closure of the fishery in Mexico (June-August). Females had more significant shell height at first maturity than males at both sites, possibly associated with fishing pressure. The information from this study is a significant contribution, but it is necessary to extend the studies to populations further south and further north.

The polyclads (Platyhelminthes: Polycladida) from the Tropical Eastern Pacific: Commented checklist and description of a new species

The polyclads (Platyhelminthes: Polycladida) from the Tropical Eastern Pacific: Commented checklist and description of a new species

Multidecadal underwater surveys reveal declines in marine turtles

AbstractMarine turtles are a group of imperiled marine megafauna particularly vulnerable to anthropogenic stressors. Most long‐term studies of marine turtles are based on nesting surveys which focus on numbers of eggs, hatchlings, and nesting females. However, we know less about long‐term abundance trends of immature and adult turtles in the marine environment. To address this data gap, we examined records from 35,000 underwater visual census (UVC) dives (1993–2019) and short‐term in‐water turtle survey data (2009–2014) at Cocos Island, Costa Rica. During UVCs, trained divemasters from UnderSea Hunter recorded observations of two species of marine turtles—green Chelonia mydas and hawksbill Eretmochelys imbricata. Our short‐term in‐water surveys revealed that most turtles at Cocos are greens, but both immature and mature greens occur at Cocos. We analyzed long‐term UVC data using a hierarchical modeling approach and we modeled a 26% decrease in the relative abundance of turtles observed on dives each year. Our model also revealed potential interactions between tiger sharks and turtles, finding that for each additional tiger shark present during a dive, the predicted relative abundance of turtles decreased by 43%. Lastly, our model suggested the influence of environmental variation on marine turtle relative abundance; a 1°C increase in sea surface temperature (SST) decreased the predicted relative abundance of turtles by 7%. Our results suggest that marine turtles are sensitive to long‐term environmental and oceanographic changes, and potentially avoid certain areas to reduce exposure to tiger sharks. Given our study area is already protected, there needs to be more focus on protecting adult turtles during their movements across the Eastern Tropical Pacific. Our work also highlights the importance of long‐term underwater surveys to monitor adult turtles.

Projected Antarctic sea ice change contributes to increased occurrence of strong El Niño

Current climate models suggested that Antarctic sea ice cover would decrease substantially under cumulative CO2 emission, but little is known whether large decrease in Antarctic sea ice can influence the occurrence of strong El Niño. Using time slice coupled and uncoupled model experiments, we show that in response to half reduction of Antarctic sea ice projected near the end of the 21st century, the frequency of strong El Niño would be increased by ~40%. It is contributed by enhanced thermocline, Ekman, and zonal advective positive feedbacks that are partly offset by enhanced thermodynamic damping. The strong warming and weakened westerly winds in the southeastern Pacific generate an anomalous Rossby wave propagating into the eastern subtropical and tropical Pacific, favoring stronger El Nino, and air-sea coupling and ocean feedbacks play a critical role in the teleconnection. Unexpectedly, compare to halved Antarctic sea ice, the ice-free Antarctic leads to a decrease in the frequency of strong El Niño, which is largely due to a substantial increase in thermodynamic damping. We also show that a large portion of the increase of strong El Niño events under greenhouse warming might be connected with Antarctic sea-ice loss, though increased greenhouse gas plays an important role.

Endolithic Algae (Ostreobium) Diversity in Porites Corals at the Western Atlantic and Tropical Eastern Pacific

ABSTRACTOstreobium comprises endolithic algae commonly seen in conjunction with scleractinian corals. In the past, it was solely recognized as a coral skeleton bioeroder. Their relationship with corals is critical because they give photosynthetic byproducts and help the coral when it loses its primary symbionts due to stress. The variety of these algae in coral species of the genus Porites in the Tropical Eastern Pacific and western Atlantic was investigated. Ostreobium samples from seven Porites species including two from the Tropical Eastern Pacific (TEP) (P. panamensis and P. lobata) and five from the Caribbean (P. furcata, P. porites, P. colonensis, P. branneri, and P. astreoides) were extracted. Published rbcL sequences from algae found within various coral species from other parts of the world were also compared. A biogeographic analysis and two methodologies, PTP (Poisson tree process) and GMYC (general mixed Yule‐coalescent), were used to delineate the different species. The findings revealed a significant degree of genetic diversity within Ostreobium, with more than 15 groups of not more than three individuals and 40 individual lineages. Its origins date back to the Ordovician, 500 Ma, and it does not appear to preserve host specificity. The sampled locations still have a wide variety of Ostreobium. Biogeographically patterns were also confirmed, making it impossible to pinpoint the precise origins of most clades. The ancestry analyses revealed convergent events for not only the emergence of Ostreobium in a few genera of local corals, but also the phenomenon occurred in genera from far‐off places.

A climate change signal in the tropical Pacific emerges from decadal variability.

The eastern tropical Pacific has defied the global warming trend. There has been a debate about whether this observed trend is forced or natural (i.e., the Interdecadal Pacific Oscillation; IPO) and this study shows that there are two patterns, one that oscillates along with the IPO, and one that is emerging since the mid-1950s, herein called the Pacific Climate Change (PCC) pattern. Here we show these have distinctive and distinguishable atmosphere-ocean signatures. While the IPO features a meridionally broad wedge-shaped SST pattern, the PCC pattern is marked by a narrow equatorial cooling band. These different SST patterns are related to distinct wind-driven ocean dynamical processes. We further show that the recent trends during the satellite era are a combination of IPO and PCC. Our findings set a path to distinguish climate change signals from internal variability through the underlying dynamics of each.

Geographic genetic variation in the Coral Hawkfish, Cirrhitichthys oxycephalus (Cirrhitidae), in relation to biogeographic barriers across the Tropical Indo-Pacific

The Tropical Indo-Pacific (TIP) includes about two thirds of the world’s tropical oceans and harbors an enormous number of marine species. The distributions of those species within the region is affected by habitat discontinuities and oceanographic features. As well as many smaller ones, the TIP contains seven large recognized biogeographic barriers that separate the Red Sea and Indian Ocean, the Indian from the Pacific Ocean, the central and eastern Pacific, the Hawaiian archipelago, the Marquesas and Easter Islands. We examined the genetic structuring of populations of Cirrhitichthys oxycephalus, a small cryptic species of reef fish, across its geographic range, which spans the longitudinal limits of the TIP. We assessed geographic variation in the mitochondrial cytb gene and the nuclear RAG1 gene, using 166 samples collected in 46 localities from the western to eastern edges of the TIP. Sequences from cytb show three well-structured groups that are separated by large genetic distances (1.58–2.96%): two in the Tropical Eastern Pacific (TEP), one at Clipperton Atoll another occupying the rest of that region and the third that ranges across the remainder of the TIP, from the central Pacific to the Red Sea and South Africa. These results indicate that the ~4,000 km wide Eastern Pacific Barrier between the central and eastern Pacific is an efficient barrier separating the two main groups. Further, the ~950 km of open ocean that isolates Clipperton Atoll from the rest of the TEP is also an effective barrier. Contrary to many other cases, various major and minor barriers from the Central Indo-Pacific to the Red Sea are not effective against dispersal by C. oxycephalus, although this species has not colonized the Hawiian islands and Easter Island. The nuclear gene partially supports the genetic structure evident in cytb, although all haplotypes are geographically mixed.

Rare occurrence of only female flowers suggests a lack of sexual reproduction and potential clonality of the seagrass Halophila baillonii Asch. on the Pacific coast of Costa Rica

Halophila baillonii Asch. is a rare seagrass species found in the Caribbean, Atlantic, and Eastern Tropical Pacific (ETP). It is listed as Vulnerable in the IUCN Red List yet very little is known about its dispersal mechanisms. For this study, samples were collected at five sites on the Pacific coast of Costa Rica. Each site was visited once between 2019 and 2021 as part of a larger molecular study. At each site, H. baillonii foliar shoots were manually collected along three transects. Presence of flowers or fruits was assessed for 1292 foliar shoots using a dissecting microscope. Additionally, sediment samples were collected to assess seed density at three of the sites. Flowering frequency was very low overall (0.7 %), flowers were only found at two of the five sites, and only female flowers were present. There were no fruits or seeds at any of the sites. This is the first report of H. baillonii flowering on the Pacific coast of Costa Rica and within the ETP. Meanwhile, H. baillonii flowers of both sexes and fruits have been found from limited sampling efforts at multiple sites in the Caribbean and Brazil. The findings of this study suggest a lack of sexual reproduction and potential clonality of the seagrass H. baillonii on the Pacific coast of Costa Rica. This supports the recent notion that H. baillonii is the first non-native seagrass in the ETP, yet more detailed field and molecular studies are needed.

Insights into Blue Whale (Balaenoptera musculus L.) Population Movements in the Galapagos Archipelago and Southeast Pacific.

The Galapagos Marine Reserve is vital for cetaceans, serving as both a stopover and residency site. However, blue whales, occasionally sighted here, exhibit poorly understood migratory behavior within the Galapagos and the broader Eastern Tropical Pacific. This study, the first to satellite tag blue whales in the Galapagos (16 tagged between 2021 and 2023), explored their behavior in relation to environmental variables like chlorophyll-a concentration, sea surface temperature (SST), and productivity. Key findings show a strong correlation between foraging behavior, high chlorophyll-a levels, productivity, and lower SSTs, indicating a preference for food-rich areas. Additionally, there is a notable association with geomorphic features like ridges, which potentially enhance food abundance. Most tagged whales stayed near the Galapagos archipelago, with higher concentrations observed around Isabela Island, which is increasingly frequented by tourist vessels, posing heightened ship strike risks. Some whales ventured into Ecuador's exclusive economic zone, while one migrated southward to Peru. The strong 2023 El Niño-Southern Oscillation event led to SST and primary production changes, likely impacting whale resource availability. Our study provides crucial insights into blue whale habitat utilization, informing adaptive management strategies to mitigate ship strike risks and address altered migration routes due to climate-driven environmental shifts.

A new species of Heterogorgia Verrill, 1868 (Octocorallia, Malacalcyonacea) from a mesophotic fishing shoal in the eastern Pacific

Heterogorgia abdita sp. nov. is a newly identified mesophotic octocoral species collected at Tigre shoal off Santa Elena Peninsula (Pacific Costa Rica). The new species is described based on distinctive morphological characters that classify it within the genus Heterogorgia. It is characterised by its conspicuous small branching colony with prominent calyces and thin branches. While its scleroma aligns with the genus, the presence of club-like spindles is notable, as these have previously been observed only in the sole Atlantic Heterogorgia species. This study enhances our understanding of the intriguing genus Heterogorgia and contributes to the knowledge of octocoral biodiversity within the Área de Conservación Guanacaste north Pacific Costa Rica and the broader marine biodiversity of the eastern tropical Pacific.

Testing the feasibility of coral nurseries in an upwelling area in the North Pacific of Costa Rica

The decline of coral reefs has increased interest in ecological restoration. Due to the scarcity of coral gardening projects in the Eastern Tropical Pacific, improving our understanding of such techniques is key. We report the results of coral gardening using the branching Pocillopora spp. and massive coral species (Pavona gigantea, Pavona clavus and Porites lobata) in an upwelling area in Costa Rica. We examined whether nursery type influenced Pocillopora spp. survival and growth, and how environmental conditions shaped restoration. We monitored the survival and growth of Pocillopora spp. fragments (n = 334) and microfragments of massive species (P. gigantea [n = 148], P. clavus [n = 37], P. lobata [n = 66]) over 11 months. Survival at the end of the gardening period was 51% for Pocillopora spp., 59% for P. clavus, 55% for P. gigantea, and 17% for P. lobata, with a decline after a cease in maintenance caused by the COVID-19 lockdown. Pocillopora spp. fragments in the floating nurseries exhibited higher growth (7.52 ± 1.98 and 6.64 ± 2.91 cm yr-1) than in the A-frame (4.16 ± 2.35 cm yr-1), which suggests the benefits of suspending fragments. For massive microfragments coral growth was 1.92-4.66 cm2 yr-1 and were affected by pigmentation loss, causing partial tissue loss and mortality. Our results point towards acclimation to local conditions, and show the need to develop site-specific cost-efficient gardening techniques for massive species, allowing for a multi-species approach to ensure long-term ecosystem recovery.

The 2023 Atlantic Hurricane Season: An Above-Normal Season despite Strong El Niño Conditions

Abstract The 2023 Atlantic hurricane season was above normal, producing 20 named storms, 7 hurricanes, 3 major hurricanes, and seasonal accumulated cyclone energy that exceeded the 1991–2020 average. Hurricane Idalia was the most damaging hurricane of the year, making landfall as a Category 3 hurricane in Florida, resulting in eight direct fatalities and 3.6 billion U.S. dollars in damage. The above-normal 2023 hurricane season occurred during a strong El Niño event. El Niño events tend to be associated with increased vertical wind shear across the Caribbean and tropical Atlantic, yet vertical wind shear during the peak hurricane season months of August–October was well below normal. The primary driver of the above-normal season was likely record warm tropical Atlantic sea surface temperatures (SSTs), which effectively counteracted some of the canonical impacts of El Niño. The extremely warm tropical Atlantic and Caribbean were associated with weaker-than-normal trade winds driven by an anomalously weak subtropical ridge, resulting in a positive wind–evaporation–SST feedback. We tested atmospheric circulation sensitivity to SSTs in both the tropical and subtropical Pacific and the Atlantic using the atmospheric component of the Community Earth System Model, version 2.3. We found that the extremely warm Atlantic was the primary driver of the reduced vertical wind shear relative to other moderate/strong El Niño events. The concentrated warmth in the eastern tropical Pacific in August–October may have contributed to increased levels of vertical wind shear than if the warming had been more evenly spread across the eastern and central tropical Pacific.

Reproductive cycle of the rock oyster Striostrea prismatica in a natural and estuarine-lagoon environment of the Mexican Central Pacific

ABSTRACT Striostrea prismatica is an oyster of commercial importance present in several locations of the Eastern Tropical Pacific. The objective of this study was to evaluate the sex ratio, gametogenic cycle, maturity stages, size at first maturity (SH50) and monthly variation in the condition index of this species over the course of an annual cycle on the coast of Colima, Mexico. The oysters found in the shallow sublittoral areas near Faro de Campos were larger and heavier than those in Cuyutlan lagoon. The sex ratio was significantly shifted towards males at both sites. The gonad showed a prolonged undifferentiated stage lasting from November to April, a spawning period limited to June through October and an extended post-spawning period from August to December. SH50 was higher in females in the lagoon as well as in the shallow sublittoral. Monthly variation in surface temperature and chlorophyll a in both study areas only correlated with the condition index of populations in the shallow sublittoral and inner lagoon in the months of lowest temperatures and highest chlorophyll a. These findings provide basic information on the reproductive cycle of rock oysters that may prove useful for the development of improved harvesting practices in the study region.

Analyzing Species Diversity in Rocky Intertidal Communities over Multiple Spatial Scales among Understudied Eastern Pacific Ecoregions

Many gaps in our theoretical understanding of the variations in the diversity and structure of intertidal communities exist for the Eastern Pacific. In order to fill some of these gaps, we censused intertidal communities and compared patterns of diversity on multiple spatial scales using several measures in alpha (α) and beta (β) diversities at twenty-one sites in a cold temperate, a warm temperate and a tropical Eastern Pacific ecoregion that were unique in terms of research effort and each with distinct geographic features. Diversity and richness on all spatial scales were compared using area curves, Hill numbers, ordination and cluster analyses, and the Hutcheson’s t-test with post hoc PERMANOVA, which revealed significant differences in diversity within and among ecoregions. Functional group and species richness and abundance were found to be highest in the cold and warm temperate ecoregions, and the functional group richness was second highest in the tropical Guayaquil ecoregion. The Bray–Curtis similarity method proved useful for determining patterns of small-scale intertidal zonation, while the Sorensen–Dice method suggested high indices of similarity in the functional group and subclass structures among all ecoregions.

Severe cold-water bleaching of a deep-water reef underscores future challenges for Mesophotic Coral Ecosystems

Elevated sea surface temperatures are causing an increase in coral bleaching events worldwide, and represent an existential threat to coral reefs. Early studies of Mesophotic Coral Ecosystems (MCEs) highlighted their potential as thermal refuges for shallow-water coral species in the face of predicted 21st century warming. However, recent genetic evidence implies that limited ecological connectivity between shallow- and deep-water coral communities inhibits their effectiveness as refugia; instead MCEs host distinct endemic communities that are ecologically significant in and of themselves. In either scenario, understanding the response of MCEs to climate change is critical given their ecological significance and widespread global distribution. Such an understanding has so far eluded the community, however, because of the challenges associated with long-term field monitoring, the stochastic nature of climatic events that drive bleaching, and the paucity of deep-water observations. Here we document the first observed cold-water bleaching of a mesophotic coral reef at Clipperton Atoll, a remote Eastern Tropical Pacific (ETP) atoll with high coral cover and a well-developed MCE. The severe bleaching (>70 % partially or fully bleached coral cover at 32 m depth) was driven by an anomalously shallow thermocline, and highlights a significant and previously unreported challenge for MCEs. Prompted by these observations, we compiled published cold-water bleaching events for the ETP, and demonstrate that the timing of past cold-water bleaching events in the ETP coincides with decadal oscillations in mean zonal wind strength and thermocline depth. The latter observation suggests any future intensification of easterly winds in the Pacific could be a significant concern for its MCEs. Our observations, in combination with recent reports of warm-water bleaching of Red Sea and Indian Ocean MCEs, highlight that 21st century MCEs in the Eastern Pacific face a two-pronged challenge: warm-water bleaching from above, and cold-water bleaching from below.

Impact of Warming Trend in Western Equatorial Pacific on Modulating the Triple‐Dip La Niña and Its Associated Teleconnection in 2020–2022

AbstractIn this study, we investigated the triple‐dip La Niña during 2020–2022 by comparing it with the previous (1973–1975 and 1998–2000) La Niña events. We found that the cold sea surface temperature (SST) in the eastern equatorial Pacific was moderate during the study period; however, the accompanying near‐surface easterly wind anomaly was unusually stronger during its lifecycle than during the previous two events. The maintenance of 2020–2022 La Niña appeared to be attributable to the strong zonal SST gradient. The strong zonal SST gradient resulted from the La‐Niña‐associated interannual SST anomaly, which was further enhanced by a warming trend in the western equatorial Pacific (165°E−160°W, 5°S–5°N) and the interdecadal oscillation of the Pacific‐Decadal‐Oscillation‐associated cold SST in the eastern tropical Pacific. The warming trend in the western equatorial Pacific, with a faster warming speed than global warming, also modified the La‐Niña‐associated Pacific–North American teleconnection to shifted eastward.

Autobiography: A 50-Year Quest for Understanding in Geoscience

Readers will be led down a random path from continental dynamics to paleoclimate. A key to understanding continental dynamics is recognizing that differences in gravitational potential energy per unit area between high and low terrain govern much of large-scale continental deformation. Removal of mantle lithosphere, not just crustal thickening, plays a crucial, but difficult-to-test, role in changes in surface elevation. Although measuring past surface heights remains a challenge, indications of such processes suggest that surface uplift associated with such removal can affect relative plate motion. Climate change, from a warmer to cooler climate, and associated changes in erosion and sedimentation introduce further complications to determining past elevations. The phenomena that led to such cooling include a number of possibilities, but I favor the emergence of islands in the Maritime continent, which transformed the Pacific Ocean from one with a warm eastern tropical Pacific, as during El Niño events, to the present-day La Niña–like background state. Teleconnections from the eastern tropical Pacific to Canada affect the duration of summers and the potential of high-latitude ice to accumulate. ▪Lateral gradients in gravitational potential energy per unit area (GPE), a force per unit length, govern large-scale continental dynamics.▪Removal of mantle lithosphere and thickening of crust raise GPE; knowledge of mean surface elevations provides a test of these processes.▪Climate change from a warmer to cooler climate and from one with less to more erosion can give the false impression of elevation change.▪Emergence of Indonesian islands, more rain over them, a stronger Walker Circulation, and cooler eastern Pacific may have led to ice ages.

Multidecadal Changes of the Seasonal Potential Predictability of Winter PNA and Associated Circulation Anomalies

Abstract Utilizing ensemble hindcast data from the Community Earth System Model (CESM) spanning the years 1900–2014, the multidecadal changes in the seasonal potential predictability of the winter Pacific–North American (PNA) teleconnection pattern and associated circulation anomalies have been investigated by using an information-based metric of relative entropy and the method of the most predictable component analysis. Results show that the seasonal potential predictability of winter PNA has significant multidecadal changes, with values much higher at the two ends of the twentieth century and much lower in between particularly in the 1930s and 1940s. The changes in the seasonal potential predictability of winter PNA are mostly reflected by the temporal evolutions of PNA rather than the location changes of active centers. Further, the changes are mostly contributed by the external forcing of El Niño–Southern Oscillation (ENSO)-related sea surface temperature anomalies in tropical central and eastern Pacific. In particular, the combined effects of lower amplitudes, reduced persistence, and a more eastward shift in warming centers lead to the reduced seasonal potential predictability of PNA and associated circulation changes in the 1930s and 1940s. Significance Statement Seasonal prediction of the winter Pacific–North American (PNA) teleconnection pattern and associated circulation anomalies is very important due to its profound climate impacts. Understanding the multidecadal fluctuations and its driving sources of the potential predictability of winter PNA and associated circulation anomalies are meaningful for skillful seasonal prediction of winter PNA and circulation anomalies as well as related climate variations. This study for the first time shows that the multidecadal fluctuations of the potential predictability of winter PNA are quite significant and the changes are mostly reflected by its temporal evolutions rather than spatial shifts of active centers. Furthermore, this study shows that the strength, persistence, and warming center locations of ENSO-related sea surface temperatures in tropical Pacific play a crucial role on the multidecadal changes of the potential predictability of winter PNA and associated circulation anomalies.

The connection of the Costa Rica Coastal Current with the West Mexican Current in the Gulf of Tehuantepec

The interconnections of some ocean currents in the Eastern Tropical Pacific are still a mystery to the scientific community. In the Mexican Pacific, the West Mexican Current (WMC) and the Costa Rica Coastal Current (CRCC) are two flows that are typically considered independent; however, some evidence of a subsurface connection when the Tehuantepec Bowl (TB) moves westward suggests otherwise. With the Hybrid Coordinated Ocean Model (HYCOM), we obtained a 19 years-run in a domain encompassing the Eastern Tropical Pacific Ocean. The numerical results were validated qualitatively and quantitatively by comparing them with remote and direct observations. We found a generally good agreement between the model results and those observations. To analyze the model results, we defined four transects perpendicularly to the coast, and surface and subsurface layers; in the long-term mean, the CRCC is disconnected from the WMC, in both layers. However, during the seasonal cycle, we found a close relationship between the general structures in thermocline topography and surface and subsurface large-scale circulation. The model results confirmed that the interconnection between the CRCC and the WMC depends on the dynamics of the TB, that on its seasonal migration moves westward, allowing the CRCC to travel along the coast further west than the Gulf of Tehuantepec (GT) and to connect with the WMC during spring at a subsurface level below the thermocline.