Early Caradoc Research Articles

Middle-Upper Ordovician conodonts from the Ffairfach and Golden Grove groups in South Wales, United Kingdom

ABSTRACT The conodont fauna of the reference succession of the regional British Llandeilian Stage of the Llanvirn Series was first described in a classical study by Rhodes more than 65 years ago using single element (form) taxonomy. Although several subsequent authors have recorded a substantial number of conodont taxa from the Llandeilo area, the present study is the first to present a modern taxonomic review of these late Darriwilian-early Sandbian faunas that include approximately 20 multielement species. Most prominent are representatives of Amorphognathus, Baltoniodus, Eoplacognathus, and Plectodina. The study faunas have their own biogeographical character. The distinctive genera Complexodus, Protopanderodus, and Pygodus, which are common in coeval Baltoscandic faunas, are not present, but the occurrence of Amorphognathus, Baltoniodus, and Eoplacognathus provides a link to age equivalent Baltoscandic faunas. The presence of abundant specimens of Plectodina and less common representatives of Erismodus and Icriodella are reminiscent of North American Midcontinent faunas. This type of faunal assemblage is in some respects similar to those of the early Caradoc Series of the Welsh Borderland. Biostratigraphically diagnostic species indicate that the Llandeilo study succession ranges from the Eoplacognathus lindstroemi Subzone of the Pygodus serra Zone to the Baltoniodus variabilis Subzone of the Amorphognathus tvaerensis Zone.

A new runner-like cyclostome bryozoan from the Bromide Formation (Sandbian, Upper Ordovician) of Oklahoma and its phylogenetic affinities

AbstractZigzagopora wigleyensis n. gen. n. sp. is an Upper Ordovician (Sandbian, early Caradoc) cyclostome bryozoan from the Arbuckle Mountains of Oklahoma, USA. It has runner-type colonies characterized by a mostly uniserial, geniculate arrangement of monomorphic zooids that bud alternately left and right, producing a zig-zag pattern of growth. This new genus has calcified interior walls and non-pseudoporous exterior walls. It is thus most likely affiliated with the paleotubuliporine family Sagenellidae, despite superficial similarities with the corynotrypid cyclostomes with which it co-occurs.

Distribution and palaeoecology of Ordovician bivalves and gastropods from Girvan, SW Scotland

ABSTRACTMolluscs from the Middle and Upper Ordovician succession of Girvan, SW Scotland are common and diverse in some localities. The mollusc fauna consists mainly of gastropods, bivalves and various univalved molluscs (mimospirids and tergomyans), along with scarcer polyplacophorans, rostroconchs and cephalopods. The present study gives an overview of the distribution and palaeoecology of bivalves, gastropods and univalved molluscs and compares them with mollusc faunas worldwide. Gastropods, mimospirids and tergomyans are present from the Darriwilian (mid Llanvirn) onwards in both siliciclastic and carbonate facies, and increase in diversity through the Sandbian (Caradoc) and into the Katian (Ashgill). Bivalves first appeared in Girvan in the late Darriwilian (early Caradoc) in deep water siliciclastic facies; where they continued to be more abundant and diverse than in equivalent carbonate facies. Molluscs are initially Laurentian in aspect, though peri-Gondwanan faunal elements occur, particularly during the Sandbian. The pattern of bivalve and gastropod diversity found in the Ordovician of Girvan generally follows that of the known global diversity for these groups.

Silicified Upper Ordovician trilobites from Pai‐Khoi, Arctic Russia

Abstract: A collection of silicified trilobites extracted from samples obtained by F. Nansen from the Upper Ordovician of Khabarovo, Pai Khoi, Russian Arctic is dominated by two new species of the proetide Lasarchopyge, a genus known hitherto only from the Argentine Precordillera. Comparison of Lasarchopyge with Scharyia reveals a range of common characters that support inclusion of both genera in the same family. The rest of the fauna comprises a metagnostid (Trinodus elspethi), an asaphid (Isotelus sp.), two remopleuridids (Remopleurides cf. caelatus and R. sp. nov. A), an aulacopleurid (Harpidella triloba), a lichid and an odontopleurid (Apianurus cf. barbatus), the last two being represented by very fragmentary material. T. elspethi, H. triloba, R. cf. caelatus and A. cf. barbatus are common to or closely resemble taxa from the Edinburg Formation, Virginia. Closely related taxa are present also in the Esbataottine Formation, NW Territories, Canada, and in the Albany Group, Girvan district, Scotland. All of these occurrences are of early Caradoc (Sandbian) age, and most lay at palaeolatitudes of 20°–30° south; all are interpreted as having been deposited on the outer shelf, which is consistent with their distribution.

Hydrodynamic control of sedimentation in the Ordovician (Arenig–Caradoc) Baltic Basin

Changes in mineralogy, geochemistry and grain size in a deep shelf succession from the Ordovician Baltic Basin are used to determine shifts in marine current direction and sea-level fluctuations. The analyses show a long-term, relatively stable interval in the Arenig–Llanvirn followed by shorter periods of perturbations and rearrangements in the early Caradoc. The Arenig deposits of the Aizpute-41 core have a high 2–10 µm terrigenous component. During the Llanvirn the proportion of this size fraction diminished. The grain size change was accompanied by mineralogical changes. In the early Caradoc the kaolinite disappeared, the muscovite content decreased, and quartz and albite increased. This and several geochemical indicators point to an input of considerable part of sediment from different sources than in the Arenig–Llanvirn. Two currents, a southwesterly for the Arenig and a westerly for the early Caradoc, are proposed. A red facies belt in the deep shelf marks the pathway of the Arenig–Llanvirn current. Smaller shifts in SiO2/Al2O3 ratio in the Aizpute-41 section are interpreted as reflecting transgressions and regressions corresponding to regional stage boundaries.

Assessing the biodiversity of Ordovician ostracods from the Argentine Precordillera

Ostracod faunas from the Lower to Middle Ordovician rocks of the Argentine Precordillera Basin (Gualcamayo and Las Aguaditas Formations) are studied. A new family, Garcianidae, is erected. One new genus,Jeanvannieria, and six species are recognized, two of which are new (Jachalipisthia bicornataandJeanvannieria bulbosa). The diversity and composition of the Precordilleran ostracods is evaluated on the basis of previous taxonomic analysis and the fauna studied here. The diversity is moderate, with a peak of 50 species during the early Caradoc. The composition of the fauna is characterized by the dominance of podocopes with a high percentage of binodicopes and a lack of palaeocopes, which is in agreement with a deep shelf environment. The carbonate slope setting of the Las Aguaditas Formation is the deepest environment yet found with Ordovician ostracods and records a relatively diverse fauna. The presence ofEctoprimitioidessuggests biogeographic affinities between the Precordillera and Laurentia. The rest of the fauna contains a high percentage of endemic genera and a mixture of genera with several affinities, Baltic, peri-Gondwanan, and Australian.

Darriwilian graptolites of the Hamburg succession (Dauphin Formation), Pennsylvania, and their geologic significance

Graptolites from the Dauphin Formation in the allochthonous Hamburg succession of the Appalachians in Pennsylvania, USA, are late Darriwilian (Da) 3 to early Da 4 age (Middle Ordovician); this age range constrains the timing of the latest depositional episode before the terrane was tectonically mobilized. These rocks were emplaced into the Martinsburg foreland basin of Laurentia during the Taconic orogeny in the early Caradoc (Late Ordovician). Nineteen taxa are described defining a narrow biostratigraphic interval. Among the characteristic fauna collected from of the Da 4 Zone are Pterograptus elegans Holm, Cryptograptus schaeferi Lapworth, Hustedograptus teretiusculus (Hisinger) ?, Haddingograptus oliveri (Bouček), Glossograptus hincksii (Hopkinson), Pseudophyllograptus angustifolius s.l. (J. Hall), and Archiclimacograptus cf. riddellensis (Harris). Tetragraptus cf. erectus Mu et al. found with the above suggests a level low in the Da 4 Zone and a limited occurrence of Bergstromograptus crawfordi (Harris) may indicate some strata within the Da 3 Zone. Proposed new forms include Pseudotrigonograptus ? ricardo sp. nov., and at least two reteograptids. Four examples of Kalpinograptus and Kalpinograptus ? may be new.

TRIARTHRINID TRILOBITES (OLENIDAE) FROM THE MIDDLE AND UPPER ORDOVICIAN, PRECORDILLERA OF ARGENTINA

Middle and Upper Ordovician Triarthrinae from the Precordillera of San Juan, Argentina, include the Whiterock/Llanvirn Porterfieldia turneri (Baldis and Pöthe, 1995) and P. acava new species, and the early Caradoc Triarthrus jachalensis (Harrington and Leanza, 1957). Each of these species is described based on silicified material, including ontogenies, from the Las Aguaditas Formation. Porterfieldia maanssonae new species from the lower member of the Gualcamayo Formation (late Arenig) at Río Gualcamayo, San Juan Province, and Río Guandacol, La Rioja Province, is closely allied to P. turneri. Two protaspid stages are present in Porterfieldia, whereas mineralized protaspid stages are apparently lacking in Triarthrus jachalensis. Triarthrus jachalensis was capable of sphaeroidal enrollment by meraspid degree 2. Precordilleran species bear closest comparison to others from Spitsbergen (Porterfieldia acava n. sp. and P. parapunctata) and Sweden/Norway (Triarthrus jachalensis and T. linnarssoni).

First Late Ordovician Paleomagnetic Pole for the Cuyania (Precordillera) Terrane of Western Argentina: a Microcontinent or a Laurentian Plateau

Time and tectonic processes involved in docking of the Argentine Precordillera (Cuyania terrane) against SW Gondwana has been a matter of much debate. A paleomagnetic study on the Early Caradoc Pavón Formation, exposed in the San Rafael block, province of Mendoza, Argentina, is presented. After detailed thermal and alternating field demagnetizations two geologically significant magnetic components were defined. A widespread post-tectonic component (A) is present in most sites of the Pavón Formation, with dual polarities, and is coincident with the characteristic remanence isolated from a Permo-Triassic rhyolitic dome intruding the sediments. Its pole position (83.7°S, 271.0°E, dp = 6.8°, dm = 9.0° N = 11 sites) falls on the Late Permian-Early Triassic South American reference poles suggesting that this component was acquired during the Choiyoi magmatic phase. A second component (B) also shows dual polarities and a positive fold test suggesting a primary origin. Unblocking temperatures and rock magnetic experiments indicate that B is carried either by hematite or magnetite at different sites. Anisotropy of magnetic susceptibility results suggest a depositional fabric and no remanence distortion due to deformation or compaction. A paleomagnetic pole computed from this remanence (PV) falls on 3.6°N, 346.4°E (dp = 2.9°, dm = 4.6° n = 22 samples). It indicates a paleolatitude around 26°S for deposition of Pavón sediments and constrains the paleogeographic evolution of Cuyania during the Ordovician, which was still at subtropical latitudes by the Early Caradoc. PV is consistent with the Laurentian Late Ordovician reference pole if Cuyania remains attached to SE Laurentia for the Early Caradoc, while it shows a significant cw rotation with no paleolatitude anomaly respect to the Gondwana reference pole when kept in its present position in SW South America. These comparisons are interpreted in three possible alternatives for the paleogeographic and tectonic setting of Cuyania in the Late Ordovician.

Biostratigraphy, taxonomic diversity and patterns of morphological evolution of Ordovician acritarchs (organic-walled microphytoplankton) from the northern Gondwana margin in relation to palaeoclimatic and palaeogeographic changes

Acritarchs, the fossilizable, resting cysts of phytoplanktonic algal protists, are the dominant component of marine organic-walled microfossils in the Palaeozoic. The majority of acritarchs show strong similarities with dinoflagellate cysts in morphological and biogeochemical features, as well as distributional patterns in the sediments. The production of these organic-walled microfossils and their distribution and survivorship in the sediments were controlled by differences in ecological tolerances and life cycle (autecology) of the planktonic parent organisms. Calculation of evolutionary rates and development of a detailed diversity curve at specific level, form the basis for discussing the influence of global palaeoenvironmental perturbations on the evolution of organic-walled microphytoplankton in northern Gondwana during latest Cambrian through Ordovician times. The potential of acritarchs for biostratigraphic correlation at the regional scale (northern Gondwana domain) is much improved by our detailed revision of distributional patterns of 245 acritarch taxa. The most important Cambro-Ordovician acritarch bio-events are short periods of diversification, which also correspond to introduction of morphological innovations, observed in latest Cambrian and earliest Tremadoc, late Tremadoc, early Arenig, basal Llanvirn, and latest Ashgill, and an important extinction phase in the early Caradoc. Overall, acritarch diversity increased from the basal Ordovician up to the middle Llanvirn, then declined in the early and middle Caradoc. During Ashgill times, the assemblages are poorly diversified at the generic level as a result of a combined effect of sea level drawdown and onset of glacial conditions, but no major extinction event is observed in connection with the end-Ordovician biotic crisis. The peak in acritarch diversity during Middle Ordovician times appears to be correlated to maximum spread of palaeogeographical assembly. Acritarch dynamics appear largely uncorrelated to second order sea-level oscillations; the primary abiotic controls on acritarch evolution were palaeogeographical and the associated palaeoceanographic changes (especially during Middle Ordovician), and the end-Ordovician palaeoclimatic shift. The acritarch fossil record provides important information on the evolution of oceanic primary producers, however, the relationships between acritarch diversity, oceanic productivity, and evolution of invertebrate animals are proving much more complex than previously thought. In particular, the hypothesis of a causal relationship between changes in acritarch diversity and metazoan evolution in the Palaeozoic is not supported by our data.

Biostratigraphy, taxonomic diversity and patterns of morphological evolution of Ordovician acritarchs (organic-walled microphytoplankton) from the northern Gondwana margin in relation to palaeoclimatic and palaeogeographic changes

Acritarchs, the fossilizable, resting cysts of phytoplanktonic algal protists, are the dominant component of marine organic-walled microfossils in the Palaeozoic. The majority of acritarchs show strong similarities with dinoflagellate cysts in morphological and biogeochemical features, as well as distributional patterns in the sediments. The production of these organic-walled microfossils and their distribution and survivorship in the sediments were controlled by differences in ecological tolerances and life cycle (autecology) of the planktonic parent organisms. Calculation of evolutionary rates and development of a detailed diversity curve at specific level, form the basis for discussing the influence of global palaeoenvironmental perturbations on the evolution of organic-walled microphytoplankton in northern Gondwana during latest Cambrian through Ordovician times. The potential of acritarchs for biostratigraphic correlation at the regional scale (northern Gondwana domain) is much improved by our detailed revision of distributional patterns of 245 acritarch taxa. The most important Cambro-Ordovician acritarch bio-events are short periods of diversification, which also correspond to introduction of morphological innovations, observed in latest Cambrian and earliest Tremadoc, late Tremadoc, early Arenig, basal Llanvirn, and latest Ashgill, and an important extinction phase in the early Caradoc. Overall, acritarch diversity increased from the basal Ordovician up to the middle Llanvirn, then declined in the early and middle Caradoc. During Ashgill times, the assemblages are poorly diversified at the generic level as a result of a combined effect of sea level drawdown and onset of glacial conditions, but no major extinction event is observed in connection with the end-Ordovician biotic crisis. The peak in acritarch diversity during Middle Ordovician times appears to be correlated to maximum spread of palaeogeographical assembly. Acritarch dynamics appear largely uncorrelated to second order sea-level oscillations; the primary abiotic controls on acritarch evolution were palaeogeographical and the associated palaeoceanographic changes (especially during Middle Ordovician), and the end-Ordovician palaeoclimatic shift. The acritarch fossil record provides important information on the evolution of oceanic primary producers, however, the relationships between acritarch diversity, oceanic productivity, and evolution of invertebrate animals are proving much more complex than previously thought. In particular, the hypothesis of a causal relationship between changes in acritarch diversity and metazoan evolution in the Palaeozoic is not supported by our data.

Huemacaspis (Trilobita, Kerfornellinae) from the Late Ordovician of the Argentine Cordillera Oriental

The trilobite genus Huemacaspis PŘIBYL and VANĚK, 1980 is redescribed from well-preserved material from the Santa Gertrudis Formation (early Caradoc), Argentine Cordillera Oriental. The new species H. gallinatoensis is described and figured. Huemacaspis is widely distributed in the southern part of the Central Andean basin during the early Caradoc and it is a key taxon for the correlation of different units from the Cordillera Oriental and Sierras Subandinas of Argentina and the Cordillera Oriental of Bolivia. This taxon is one of the few records of trilobites of early Caradoc age found in the shallow shelf environments of southwestern Gondwana. Huemacaspis is accommodated within the Kerfornellinae, a subfamily otherwise known from southern Central Europe and northern Africa. The trilobite fauna from the early Caradoc of the Central Andean basin suggests a faunal exchange with the latter regions indicating the persistence of a broadly similar biogeographic pattern to that documented in earlier Ordovician times.

Implications of Gondwana glaciations in the Baltic late Ordovician and Silurian and a carbon isotopic test of environmental cyclicity

Abstract Four glaciations – the extensively studied Hirnantian episode and three in the Llandovery-earliest Wenlock –are well established on the Gondwana palaeocontinent. New data [Hamoumi, 1999] shift the beginning of the glacial epoch to the early Caradoc when Baltica was moving from middle to low latitudes of the southern Hemisphere. Despite the rather considerable distance between polar areas of Gondwana and subtropical Baltica all four glacial events are reflected via global climatic mechanisms in the East Baltic. It is generally accepted that glaciations are marked by positive excursions of δ18O and δ13C values caused by increase of the polar ice caps, bioproduction and decrease of oceanic water temperature, etc. Based on these relationships, the Gondwana glacial events are correlated with coeval isotopic shifts established in sections of Baltica. In addition, agreement of the oceanic processes and corresponding carbon isotopic trends predicted by Jeppsson [1990] to real measured values is analysed. The following positive δ13C excursions are recorded in the Baltic area (peak values in parentheses) : middle Caradoc (2.2 ‰), early Ashgill (2.5 ‰), Hirnantian (6 ‰), early Aeronian (3.7 ‰), early Telychian (2.7 ‰), early Wenlock (5.2 ‰). Most of these shifts correlate well with glacio-eustatic sea level lowstands and biodiversity changes, as shown by the most extensive Oandu crisis in the Caradoc, Hirnantian mass extinction and the Wenlock Ireviken Event. Analysis of data allows the following conclusions : (1) all four Gondwana glaciations identified by tillites, microconglomeratic clays, etc. and dated biostratigraphically are recognised in the Baltic area through clear positive carbon isotope excursions at the same levels; (2) three smaller carbon isotope excursions in the Caradoc and Ashgill together with algal abundance data suggest the presence of several colder climate episodes during the late Ordovician. This may support the idea of the earlier onset of the glacial epoch on Gondwana; however, correct biostratigraphic dating of supposed glacial sediments is required; (3) the carbon isotopic testing of the oceanic model by Jeppsson reveals too many contradictions between model predictions and measured values. This means that the environmental background of isotopic events and relationships with oceanic events should be revised ; (4) for delimitation of the climatic –oceanic episodes, a more general marker identifying environmental change via a basinal approach seems useful. For this purpose lithological, geochemical or palaeontological criteria can be used.

Ordovician palaeogeography with new palaeomagnetic data from the Montagne Noire (Southern France)

A joint palaeomagnetic and 40Ar/39Ar study has been performed on two olistolithic blocks from the Cabrières Wildflysch in the Montagne Noire region of the Massif Central in France. There, andesitic volcanic and volcaniclastic rocks of Llanvirn–Early Caradoc age (ca 470–458 Ma) occur. Despite extensive secondary alteration, destruction of the dominant magnetic mineral phase and 40Ar/39Ar whole rock experiments that demonstrate that the volcanic rocks suffered significant argon loss, a positive fold test and the presence of dual polarities suggest that a primary, Ordovician magnetisation has mostly survived. This is one of the few documented cases where the argon system was substantially reset whilst a subordinate set of small, relatively unaltered magnetite grains, probably hosted in silicates, still carry the original, in this case Ordovician, remanence.The new data show that the Montagne Noire region was located at high southerly latitudes (68° +17/-15) during the Mid-Ordovician. This latitude represents the location for NW Gondwana of which the Massif Central was a part. Palaeomagnetic data from all the Central European massifs and terranes demonstrate a close link to the Gondwana Margin during the Lower and Middle Ordovician.

Quantifying paleogeography using biogeography: a test case for the Ordovician and Silurian of Avalonia based on brachiopods and trilobites

Despite substantial advances in plate tectonic modeling in the last three decades, the postulated position of terranes in the Paleozoic has seldom been validated by faunal data. Fewer studies still have attempted a quantitative approach to distance based on explicit data sets. As a test case, we examine the position of Avalonia in the Ordovician (Arenig, Llanvirn, early Caradoc, and Ashgill) to mid-Silurian (Wenlock) with respect to Laurentia, Baltica, and West Gondwana. Using synoptic lists of 623 trilobite genera and 622 brachiopod genera for these four plates, summarized as Venn diagrams, we have devised proportional indices of mean endemism (ME, normalized by individual plate faunas to eliminate area biogeographic effects) and complementarity (C) for objective paleobiogeographic comparisons. These can discriminate the relative position of Avalonia by assessing the optimal arrangement of inter-centroid distances (measured as great circles) between relevant pairs of continental masses. The proportional indices are used to estimate the “goodness-of-fit” of the faunal data to two widely used dynamic plate tectonic models for these time slices, those of Smith and Rush (1998) and Ross and Scotese (1997). Our faunal data are more consistent with the latter model, which we use to suggest relationships between faunal indices for the five time slices and new rescaled inter-centroid distances between all six plate pairs. We have examined linear and exponential models in relation to continental separation for these indices. For our generic data, the linear model fits distinctly better overall. The fits of indices generated by using independent trilobite and brachiopod lists are mostly similar to each other at each time slice and for a given plate, reflecting a common biogeographic signal; however, the indices vary across the time slices. Combining groups into the same matrix in a “total evidence” analysis performs better still as a measure of distance for mean endemism in the “Scotese” plate model. Four-plate mean endemism performs much better than complementarity as an indicator of pairwise distance for either plate model in the test case.

Chitinozoa and Nd isotope stratigraphy of the Ordovician rocks in the Ebbe Anticline, NW Germany

Abstract Strongly tectonized Ordovician rocks appear in the Ebbe Anticline (Rheinisches Schiefergebirge), West Germany. These fine-grained detrital rocks of the Herscheider Schichten are divided into the Plettenberger Bänderschiefer. Unterer (Kiesberter) Tonschiefer, (Rahlenberger) Grauwackenschiefer, and the Oberer (Solinger) Tonschiefer. The scope of this investigation was to improve the dating of the entire Ordovician succession. but especially the Oberer (Solinger) Tonschiefer. We used chitinozoans. Palaeozoic microfossils of high biostratigraphic value, and Nd isotopes, which previously have been used for correlation and terrane affinity analysis. Chitinozoan preservation is poor, but some taxa could be confidentially identified to the species level. The ɛNd(t) values obtained from the Ordovician succession range from -8.0 to -9.2. Joint evaluation of chitinozoan and Nd isotope data together with previously known age-ranges suggest the following ages for the Herscheider Schichten: Plettenberger Bänderschiefer (early Abereiddian, earliest Llanvirn), Unterer (Kiesberter) Tonschiefer (early to mid Abereiddian, early Llanvirn). (Rahlenberger) Grauwackenschiefer (Aurelucian, earliest Caradoc), and Oberer (Solinger) Tonschiefer (late Caradoc). The Ebbe ɛNd(t) values are most readily compared with ɛNd(t) values from Avalonia. and we therefore support the inclusion of the Ordovician rocks of the Ebbe Anticline in that palaeocontinent.

Timing of the Avalonia-Baltica plate convergence as inferred from palaeogeographic and stratigraphic data of chitinozoan assemblages in west Pomerania, northern Poland

Abstract Tectonically disturbed Ordovician rocks penetrated by deep drillholes in Pomerania, NW Poland (Koszalin-Chojnice Zone) belong to the Heligoland-Pomerania Deformation Belt. Earlier data demonstrate that the Avalonia-Baltica collision occurred in Late Ordovician times, but in Pomerania, the timing of convergence has not been ascertained, and it is uncertain if the rocks underneath the Koszalin-Chojnice Zone belong to Avalonia or Baltica. Data from chitinozoans, organic-walled Palaeozoic microfossils with applications in biostratigraphy and palaeobiogeography, were assessed from ten boreholes (Brda 2; Brda 3; Chojnice 5; Karsina 1; Kościernica 1; Nowa Wieś 1; Okunino 1; Sarbinowo 1; Skibno 1; Wyszebórz 1) to address these problems. The results improve the biostratigraphy of the cores and demonstrate that the youngest Ordovician rocks are of a Burrellian (early mid Caradoc) to Cheneyan (late mid Caradoc) age. Because these rocks are interpreted as forming part of the deformation belt, the obtained ages put a lower age limit on the initiation of foreland basin sedimentation on the foreland of the orogeny, i.e. the Baltic platform. Quantitative comparison of chitinozoan assemblages demonstrates a high level of similarity between Pomerania and Avalonia. Together, Pomerania and Avalonia show greater similarity to Baltoscandia than to North Gondwana, supporting the idea that the Tornquist Ocean had narrowed significantly in early Caradoc times.

Long-term effect of the Kärdla crater (Hiiumaa, Estonia) on Late Ordovician carbonate sedimentation

Abstract Kardla impact crater, 4 km in diameter, was formed in the early Caradoc (Late Ordovician) in the Baltoscandian shelf sea, a sediment-starved temperate-water carbonate basin. The Upper Ordovician post-impact sediments that cover the Kardla crater are 15–275 m thick. The crater rimwall is composed of three separate basement rises that formed islands or shoals in the Late Ordovician shelf sea and influenced the sedimentation in the surrounding area. The rises provide the means to study the shallow-water sedimentary environments and sea-level history. This may be one of the few cases, where shore environments of the Baltoscandian Ordovician palaeobasin are preserved in the geological record. Lithofacies distribution was studied in the crater area from data of about 100 drillcores. Grain size distribution and composition of the non-carbonate material were analyzed in six core sections in order to reconstruct the history of crater erosion. The post-impact marine deposits lie on a partly redeposited and mixed ejecta layer derived from the Cambrian to Lower Ordovician silici-clastic deposits in the target area. Beds of marine carbonate sediments of Caradoc age, wackestones and mudstones, on-lap on the slope of the crater rim. A shift of sedimentation towards the crater rim during the Caradoc is evidence of a relative sea-level rise, which was a combination of early Caradoc eustatic rise, crater area subsidence, and compaction of pre-impact and impact sediments. The occurrences of coarse sand and gravel in specific stratigraphic intervals reflect the intensity of wave action and sea-level changes in the area. A detailed curve of sea-level changes during Late Ordovician has been constructed and different post-impact sedimentation models are presented for the crater area. The sedimentation pattern changed during the Caradoc as a result of sea-level rise and climatic change. The synchronous appearance of echinoderm bioclastic accumulations in shoal areas in Estonian mainland and in the Kardla rim dates the climatic change in the basin area as late Keila (middle Caradoc) time. Assuming that the Caradoc lasted about 11 million years, infilling of the Kardla crater required about 7 million years, and influence of the crater structure on sedimentation was obvious even 10 million years after the impact event. The tectonic stability of the region, the low rate of sedimentation, and long lasting differential compaction of sediments in the crater area account for this extraordinary long influence of crater. The Kardla crater is an example of the astrobleme, formed in ancient shelf sea, which may be used to reconstruct basin history of post-impact times.

SYSTEMATICS OF SHUMARDIIDAE (TRILOBITA), WITH NEW SPECIES FROM THE ORDOVICIAN OF ARGENTINA

New shumardiid trilobites from Arenig to Early Caradoc deposits of the Argentine Precordillera include species of a revised Kweichowilla (K. salasae, K. sterrenae, and K. piojensis) and the new genus Changchowilla (C. sanjuanina and C. gracielae, and C.? carrerai). Conophrys fabiani new species occurs in the Early Tremadoc of the western Puna region, northwestern Argentina. Leoforteyia new genus is based on L. hintzei new species from the Ibexian of Utah. Published taxa recognized as new species are Conophrys rushtoni, Conophrys wrighti, Leoforteyia ludvigseni and Shumardia whittingtoni. Phylogenetic analysis suggests the differentiation of Shumardia Billings, 1862; Kweichowilla Chang, 1964; Changchowilla new genus; and Leoforteyia new genus as separate taxa and indicates that Conophrys Callaway, 1877, is a grade group. Silicified material provides a nearly complete growth series for Kweichowilla salasae, including the first protaspides known with confidence for the Shumardiidae. Additionally, ontogenetic material for K. sterrenae, C. sanjuanina, C. riojana (Benedetto and Cañas) and C.? carrerai is described.Leoforteyia is restricted to the Ibexian-Whiterockian of western Laurentia; species of Shumardia are widespread in the Arenig to Early Ashgill of western Europe, Australia, Laurentia, China and Russia. Changchowilla is recorded in the mid-Late Arenig-Llanvirn of the Precordillera and in the Llanvirn-middle Caradoc of China. Argentine species of Kweichowilla are closely allied to Australian species, suggesting biogeographic affinities of the Precordillera to the eastern margin of Gondwana by the Llanvirn and Caradoc.Functional morphology, occurrence in fine grained sediments, and association mainly with atheloptic and pelagic trilobites suggest that shumardiids were deposit-feeders that inhabited low light levels in low energy settings, with a limited ability to burrow in soft sediments.

Assessing trilobite biodiversity change in the Ordovician of the British Isles

AbstractThe Ordovician Period saw the most sustained, steep rise in marine biodiversity in the history of life on Earth and set the ecological pattern for the rest of the Palaeozoic Era. The long history of research, wide variety of depositional settings and juxtaposition of terranes with very different palaeobiogeographical histories makes the Ordovician of the British Isles an excellent laboratory in which to study this change. A database approach to analysis of trilobite biodiversity change is described using a simple compilation of stratigraphical ranges of 663 species of Avalonian trilobites, and a much more sophisticated information‐rich relational database based on the fundamental information of the fossil record–species at localities. The latter includes 2001 occurrence records of 617 trilobite species at 508 localities in the Welsh Basin. The first ever species‐level diversity curves for the whole Ordovician of an entire region are presented. Both databases reveal an overall increase in species‐ and genus‐level diversity through the Ordovician. Random resampling tests and environmental information are used to remove sampling effects from the Welsh Basin diversity curves. The main features of trilobite biodiversity change in Avalonia are the following: a late Arenig–early Llanvirn increase, during which time the highest species‐to‐genus ratios occur, is contemporaneous with a rise in global diversity and rifting of Avalonia from Gondwana; a late Abereiddian dip is followed by recovery during the Llandeilian–early Caradoc; decline during the late Caradoc–early Ashgill is at least partly attributable to lack of preserved rock and restriction of most of the preserved faunas to deep‐water environments. The greatest diversity occurs in the palaeoenvironmentally differentiated Rawtheyan and is followed by the diversity crash seen in the Hirnantian throughout the world. Copyright © 2001 John Wiley & Sons, Ltd.