Dry Ridges Research Articles

Modelling spatio‐temporal soil moisture dynamics in mountain tundra

AbstractSoil moisture has a fundamental influence on the processes and functions of tundra ecosystems. Yet, the local dynamics of soil moisture are often ignored, due to the lack of fine resolution, spatially extensive data. In this study, we modelled soil moisture with two mechanistic models, SpaFHy (a catchment‐scale hydrological model) and JSBACH (a global land surface model), and examined the results in comparison with extensive growing‐season field measurements over a mountain tundra area in northwestern Finland. Our results show that soil moisture varies considerably in the study area and this variation creates a mosaic of moisture conditions, ranging from dry ridges (growing season average 12 VWC%, Volumetric Water Content) to water‐logged mires (65 VWC%). The models, particularly SpaFHy, simulated temporal soil moisture dynamics reasonably well in parts of the landscape, but both underestimated the range of variation spatially and temporally. Soil properties and topography were important drivers of spatial variation in soil moisture dynamics. By testing the applicability of two mechanistic models to predict fine‐scale spatial and temporal variability in soil moisture, this study paves the way towards understanding the functioning of tundra ecosystems under climate change.

Evaluation of Different Drainage Systems and Sowing Dates for Improving Wheat Productivity on Vertisols of Central Highlands of Ethiopia

Water logging is a challenge to crop productivity under the rain fed system on Vertisols of central highlands of Ethiopia. Using appropriate drainage system and manipulating planting dates can reduce the effect of water logging and improve productivity of Vertisols of the area. The current field experiment was conducted to evaluate four different drainage systems ( Dry planting, Ridge &Furrow (RF), Broad Bed Maker( BBM) and Flat bed) and five sowing dates (Mid may, Early June, Mid-late June, Mid July and Late August ) in improving productivity of wheat crop on vertisol of central highlands of Ethiopia under rain fed condition during the main cropping season. The treatments of the experiment were the systematic combination of four drainage systems and five planting dates. The experiment was conducted at three locations (Ginchi on station, Sheno and Sodo dache on farm) on Vertisols of central highlands of Ethiopia. Analysis of variance indicated that some of wheat Yield related parameters (Plant height, spike length, biomass yield and grain yield ) tasted were significantly (p<0.05) affected by different drainage systems and sowing dates. According to this research trial, Out of seven treatments tasted T 2 (Early June + Dry plating) provide the highest grain yield 2214.7Kg/ha, 2230kg/ha and 8253kg/ha was recorded at Ginchi, sodo dache and sheno sites respectively. As vertisols in Ethiopia exist in different agro-climatic conditions in the country and show a considerable variability the research should be repeated and be supported with the long term local weather and soil data in order to determine drainage systems and sowing dates for a specific locations of vertisol areas. Keywords: Vertislol, Drainage system, planting date, Yield, Wheat DOI: 10.7176/JNSR/11-19-05 Publication date: October 31 st 2020

Geophysical Contribution to the Determination of the Limit between Laayoune and Foum El Oued Aquifers, Origin of Supply Sources of Oued Sakia El Hamra, Laayoune Province, Morocco

Laayoune and the Foum El Oued aquifers are in hydraulic communication only at the level of Oued Saguia El Hamra. The present study has accordingly made use of all the hydrogeological, hydrological, geological and geophysical data that preceded the watershed of Oued Saguia El Hamra in its downstream part. These data are by no means omplementary with the objective of having a better understanding of the boundary line between Laayoune and Foum El Oued aquifers and the origin of feeding the sources of Oued Saguia El Hamra. This study will focus only on the previous geophysical studies where a reinterpretation of electrical soundings has proved useful as a result of the recent well-logging results. It makes it possible to highlight the presence of a significant rise in the truncated marly substratum of Oued Saguia El Hamra and depressions (left and right banks) which could correspond to stream channels or depressed areas. At the level of the Wadi bed, there has been a regular immersion of the conductive level roofs from east to west towards Foum El Oued favoring the flow of wastewater from the zone and spraying the brackish water sources towards the groundwater of Foum El Oued. In the light of the reinterpretation of electric polls, plus as well as the geophysical surveys by electrical tomography and high definition made at the right and left banks of the Oueed Saguia El-Hamra, it was possible to verify the existence of dry ridge separating the two webs of Laayoune and Foum El Oued and stream channels or depressed areas of the left bank for drainage of brackish water to sources located along Oued Saguia El Hamra. The true resistivity models tomography profiles confirm the presence of the backbone at the left and right banks and the graben of the left bank for underground water drainage of the web Laayoune to sources welling the river Saguia El Hamra. They reveal the presence of a quaternary plio-cover (coastal platform Moghrebian) as being heterogeneous and affected by many electrical discontinuities, particularly in the level resistant R2 (coquina sandstone). These discontinuities could correspond to lateral changes in facies and/or synsedimental faults compartmentalizing the plio-quaternary formations into a system of Horsts and Grabens that relies on the whole (D1, Cs) attributed to formations from the Miocene to the Upper Cretaceous. The contours of the map show the distribution of the three families of electrical soundings A, B and C limited by two main electrical discontinuities D and M. East of discontinuity million, this map reflects the look of the wall of the Pliocene-Quaternary resting on the impermeable upper Cretaceous floor. To the west of this electrical discontinuity M, the map reflects the behavior of the roof of marl deposits of Miocene age.

Forty-Eight Years of Forest Succession: Tree Species Change across Four Forest Types in Mid-Missouri

In the central and eastern United States, many forest ecosystems have undergone recent shifts in composition and structure that may conflict with contemporary management objectives. Long-term forest inventory data were used to determine patterns of forest succession over a 48-year period for four forest types in mid-Missouri: bottomlands, dry ridge and slope, glade-like, and mesic slopes. All forest types increased in stand basal area and overstory quadratic mean diameter through time, with concomitant decreases in the number of midstory trees. Sugar maple (Acer saccharum Marshall) increased in importance value on dry ridge and slope and mesic slope forest types, largely due to the accumulation of trees in smaller diameter classes. White oak (Quercus alba L.) increased in overstory basal area in dry ridge and slope plots through the duration of the study, whereas black oak (Quercus velutina Lam.) and Shumard oak (Quercus shumardii Buckley) decreased in overstory density and basal area through time. Oak stems were nearly absent from the midstory across forest types in the recent sampling, suggesting future challenges for maintaining oak-dominated canopies following attrition of canopy trees through time on upland forest types. In glade-like plots, eastern redcedar (Juniperus virginiana L.) increased in both overstory density and basal area through time, and Shumard oak decreased in density. The importance value of chinkapin oak (Quercus muehlenbergii Engelm.) in the overstory decreased through time in glade-like plots, largely due to the increase in density of eastern redcedar rather than the loss of chinkapin oak from the overstory. The patterns of succession in this forest landscape of mid-Missouri suggest that forest management may be needed to address two common contemporary concerns: (1) the need for increasing oak advance reproduction and recruitment to maintain oak as a canopy species; and (2) reducing eastern redcedar encroachment for glade restoration and management.

Structural and compositional shifts in forests undergoing mesophication in the Wayne National Forest, southeastern Ohio

Mesophication refers to the positive feedback cycle that occurs when fire suppression causes compositional shifts from oak (Quercus spp.) to other, primarily mesophytic, species. Because mesophytic species tend to be associated with relatively high-moisture sites, we sought to understand the extent to which physiographic and edaphic factors influence this process by examining forest successional dynamics over a period of 22 years. In 2016, we resampled a network of permanent plots first sampled in 1994 that were stratified across different landscape positions (e.g., north-facing slopes vs. south-facing slopes) in the Athens Unit of the Wayne National Forest in southeastern Ohio. By studying the changes in forest structure and composition relative to ecological land type, our results suggest little, if any, resistance to mesophication on sites with low moisture availability. For example, the relative density of red maple (Acer rubrum) large saplings on dry ridges increased 25%, while zero oaks were recorded in the large sapling layer on dry ridges in 2016. However, we did observe that American beech (Fagus grandifolia) is becoming substantially more abundant in the understory and may be more influential than red maple in determining the future species composition of these forests. American beech is expected to respond differently than red maple to future scenarios under predicted climate change, and is subject to a suite of novel pathogens that have broad implications for forest management in the region.

Soil seed bank under variegated thistle does not explain thistle dominance

Variegated thistle (Silybum marianum) is a large, spiny annual that often forms dense monospecific communities on dry ridges and sunny hillslopes. The owner of a typical Poverty Bay hill-country farm with persistent variegated-thistle infestations reported that winter applications of herbicide were ineffectual in the long term as more variegated thistles simply recolonised the sprayed sites. An absence of preferred species, particularly perennial ryegrass (Lolium perenne) and legumes (Lotus and Trifolium spp.), in the soil seed bank under dense thistle populations may explain the persistence of these monospecific populations. To test this hypothesis, soil samples were collected from a dense and sparse variegated-thistle population in each of seven paddocks and incubated in a glasshouse. Emerged seedlings were identified and counted. The incubation was repeated three times. Total soil seed numbers were similar under both the dense and sparse populations with similar numbers of preferred legumes under both. However, there were significantly more perennial ryegrass seeds under the dense variegated-thistle populations compared with the sparse ones. Domination of thistles in densely infested patches was not due to lack of preferred species, or indeed other weed species, in the soil seed bank.

Spatial modeling and inventories for prioritizing investment into oak-hickory restoration

Oak (Quercus spp.) and hickory (Carya spp.) forests in the eastern United States provide a host of ecosystem services as their mast are prized by wildlife, the timber is a valued commodity, and they are generally more tolerant of extreme weather events under a changing climate. They are, however, undergoing a severe decline in prominence throughout the region, yielding to more mesic and shade-tolerant species, largely red maple (Acer rubrum). Two decades of research in Ohio have shown that silviculture and/or natural disturbances that reduce understory shade during seedling establishment and early growth, followed by canopy opening and competition management through prescribed fire and partial cutting, can encourage oak and hickory regeneration, most successfully on drier ridges and south- and southwest-facing slopes. We employed an ecological classification and mapping approach to prioritize areas across a 17-county region (∼22,000 km2) that may be more receptive, and thus more cost effective, to successful oak regeneration following silvicultural treatment. The ecomapping effort was comprised of two parts; a GIS model of the terrain, and a stand inventory of current vegetation condition coupled with the SILVAH decision-support system to recommend needed silvicultural treatments. The GIS model is based primarily on topography as vegetation patterns in the project area are largely driven by landscape position and soil moisture regimes. It uses transformed aspect, slope angle, topographic position index, and slope position as inputs to define six classes of landtype phases: ridge, southwest upper slopes, southwest lower slopes, northeast upper slopes, northeast lower slopes, and bottomland. The first three and following two classes, respectively, were hierarchically nested to form Dry Oak Forest and Dry-mesic Mixed Oak Hardwood Forest classes at the landtype level. Dry Oak Forests require the least silvicultural intervention to sustain or restore oak, while the other two landtypes normally require serious intervention to sustain oak into the future. To determine whether sufficient stocking is present for adequate regeneration, we use forest inventory data to represent current vegetation conditions including both overstory and understory stocking. Overall, these tools allow managers to identify ‘zones of investment’, i.e., those stands with the bulk of the area in the Dry Oak Forest landtype and with some level of advance oak regeneration, which will have a greater likelihood of growing into oak-dominated stands with minimal investment of scarce funding resources.

4000 Years of Changing Wetness in a Permafrost Polygon Peatland (Kytalyk, NE Siberia): A Comparative High‐Resolution Multi‐Proxy Study

AbstractIce‐wedge polygon mires feature a micro‐relief of dry ridges, shallow wet depressions, deeper wet troughs and transitional sites, resulting in a local mosaic of vegetation. The correct recognition of these landscape elements in palaeoecological studies of peat sections requires insight about the suitability of proxies and their potential for palaeoecological reconstruction in order to reconstruct vegetation and wetness patterns as well as dynamics. This paper analyses a 105.5 cm long peat section with a base dating to about 4000 cal yr BP from an ice‐wedge polygon mire near Kytalyk (NE Siberia). Pollen, macrofossils, testate amoebae, geochemistry and sediment properties were analysed in order to compare the suitability of these proxies to reconstruct past surface wetness. The proxies show similar wetness trends. Pollen and geochemistry data did not always permit wetness reconstruction, the former because many pollen types do not allow the identification of taxa at a low taxonomic resolution, the latter because later taphonomic processes modify chemical variables in deeper peat layers. Macrofossils provided the most detailed wetness reconstruction, because they could be identified to genera or species, for which the moisture requirements are accurately known from their present‐day distribution in ice‐wedge polygons. All proxies, except geochemistry, show an obvious change from wet to dry conditions at around 20 cm depth. However, as the proxies sometimes show contradictory results, a multi‐proxy approach is preferable over a single proxy interpretation as it allows the reconstruction of environmental development in a broader palaeoecological context. Copyright © 2016 John Wiley &amp; Sons, Ltd.

Spatio-temporal sensitivity of MODIS land surface temperature anomalies indicates high potential for large-scale land cover change detection in Arctic permafrost landscapes

The accelerated warming of the Arctic climate may alter the local and regional surface energy balances, for which changing land surface temperatures (LSTs) are a key indicator. Modeling current and anticipated changes in the surface energy balance requires an understanding of the spatio-temporal interactions between LSTs and land cover, both of which can be monitored globally by measurements from space. This paper investigates the accuracy of the MODIS LST/Emissivity Daily L3 Global 1km V005 product and its spatio-temporal sensitivity to land surface properties in a Canadian High Arctic permafrost landscape. The land cover ranged from fully vegetated wet sedge tundra to barren rock. MODIS LSTs were compared with in situ radiometer measurements from wet tundra areas collected over a 2-year period from July 2008 to July 2010 including both summer and winter conditions. The accuracy of the MODIS LSTs was −1.1°C with a root mean square error of 3.9°C over the entire observation period. Agreement was lowest during the freeze-back periods where MODIS LST showed a cold bias likely due to the overrepresentation of clear-sky conditions. A multi-year analysis of LST spatial anomalies, i.e., the difference between MODIS LSTs and the MODIS LST regional mean, revealed a robust spatio-temporal pattern. Highest variability in LST anomalies was found during freeze-up and thaw periods as well as for open water surface in early summer due to the presence or absence of snow or ice. The summer anomaly pattern was similar for all three years despite strong differences in precipitation, air temperature and net radiation. Summer periods with regional mean LSTs above 5.0°C showed the greatest spatial diversity with four distinct 2.0°C classes. Summer anomalies ranged from −4.5°C to 2.6°C with an average standard deviation of 1.8°C. Dry ridge areas heated up the most, while wetland areas and dry areas of sparsely vegetated bedrock with a high albedo remained coolest. The observed summer LST anomalies can be used as a baseline against which to evaluate both past and future changes in land surface properties that relate to the surface energy balance. Summer anomaly classes mainly reflected a combination of albedo and surface wetness. The potential to use this tool to monitor surface drying and wetting in the Arctic should therefore be further explored. A multi-sensor approach combining thermal satellite measurements with optical and radar imagery promises to be an effective tool for a dynamic, process-based ecosystem monitoring scheme.

Carbon accumulation in a permafrost polygon peatland: steady long-term rates in spite of shifts between dry and wet conditions.

Ice-wedge polygon peatlands contain a substantial part of the carbon stored in permafrost soils. However, little is known about their long-term carbon accumulation rates (CAR) in relation to shifts in vegetation and climate. We collected four peat profiles from one single polygon in NE Yakutia and cut them into contiguous 0.5 cm slices. Pollen density interpolation between AMS (14)C dated levels provided the time span contained in each of the sample slices, which--in combination with the volumetric carbon content--allowed for the reconstruction of CAR over decadal and centennial timescales. Vegetation representing dry palaeo-ridges and wet depressions was reconstructed with detailed micro- and macrofossil analysis. We found repeated shifts between wet and dry conditions during the past millennium. Dry ridges with associated permafrost growth originated during phases of (relatively) warm summer temperature and collapsed during relatively cold phases, illustrating the important role of vegetation and peat as intermediaries between ambient air temperature and the permafrost. The average long-term CAR across the four profiles was 10.6 ± 5.5 g C m(-2) yr(-1). Time-weighted mean CAR did not differ significantly between wet depression and dry ridge/hummock phases (10.6 ± 5.2 g C m(-2) yr(-1) and 10.3 ± 5.7 g C m(-2) yr(-1), respectively). Although we observed increased CAR in relation to warm shifts, we also found changes in the opposite direction and the highest CAR actually occurred during the Little Ice Age. In fact, CAR rather seems to be governed by strong internal feedback mechanisms and has roughly remained stable on centennial time scales. The absence of significant differences in CAR between dry ridge and wet depression phases suggests that recent warming and associated expansion of shrubs will not affect long-term rates of carbon burial in ice-wedge polygon peatlands.

Late Holocene vegetation, climate, and land-use impacts on carbon dynamics in the Florida Everglades

Tropical and subtropical peatlands are considered a significant carbon sink. The Florida Everglades includes 6000-km2 of peat-accumulating wetland; however, detailed carbon dynamics from different environments within the Everglades have not been extensively studied or compared. Here we present carbon accumulation rates from 13 cores and 4 different environments, including sawgrass ridges and sloughs, tree islands, and marl prairies, whose hydroperiods and vegetation communities differ. We find that the lowest rates of C accumulation occur in sloughs in the southern Everglades. The highest rates are found where hydroperiods are generally shorter, including near-tails of tree islands and drier ridges. Long-term average rates of 100 to >200 g C m−2 yr−1 are as high, and in some cases, higher than rates recorded from the tropics and 10–20 times higher than boreal averages. C accumulation rates were impacted by both the Medieval Climate Anomaly and the Little Ice Age, but the largest impacts to C accumulation rates over the Holocene record have been the anthropogenic changes associated with expansion of agriculture and construction of canals and levees to control movement of surface water. Water management practices in the 20th century have altered the natural hydroperiods and fire regimes of the Everglades. The Florida Everglades as a whole has acted as a significant carbon sink over the mid- to late-Holocene, but reduction of the spatial extent of the original wetland area, as well as the alteration of natural hydrology in the late 19th and 20th centuries, have significantly reduced the carbon sink capacity of this subtropical wetland.

Tetraploids do not form cushions: association of ploidy level, growth form and ecology in the High Arctic Saxifraga oppositifolia L. s. lat. (Saxifragaceae) in Svalbard

Saxifraga oppositifolia L. is a common circumpolar plant species that displays considerable morphological and genetic variation throughout its range. It is mainly diploid, but tetraploids are reported from several regions. The growth form varies from prostate to cushion-shaped, and the plant thrives in wet snow beds as well as on dry ridges. This variation has triggered the curiosity of many researchers, but as yet, no one has explained the observed morphological variation using ecological and/or genetic factors. However, the ploidy level has rarely been taken into account. This is the first study that demonstrates a significant correlation between ploidy level, ecology and growth form in S. oppositifolia . We successfully analysed 193 individuals of S. oppositifolia from 15 locations in Svalbard to investigate possible relationships among growth forms (prostrate, intermediate and cushion), ecological factors (vegetation and soil characteristics) and ploidy level. Results from flow cytometry reported 106 diploids, eight triploids and 79 tetraploids. Tetraploids almost exclusively showed prostrate growth, while the diploids displayed all three growth forms, evidence that growth form is at least partly genetically determined. Our analyses of environmental and vegetation data in relation to ploidy level indicated overlapping niches, but the tetraploids showed a narrower niche, and one shifted towards more benign habitats characterized by higher pH, higher soil temperatures and higher cover of vascular plants. The latter may suggest that tetraploids are slightly better competitors, but less hardy. Thus, autopolyploidy in S. oppositifolia has expanded the ecological amplitude of this species complex. Keywords: Autopolyploidy; flow cytometry; morphotypes; habitat segregation; purple saxifrage (Published: 6 June 2013) To access the supplementary material for this article, please see Supplementary files in the column to the right (under Article Tools) Citation: Polar Research 2013, 32 , 20071, http://dx.doi.org/10.3402/polar.v32i0.20071

Successional trends and apparent Acer saccharum regeneration failure in an oak-hickory forest in central Missouri, USA

Permanent vegetation plots established in 1968 at the Baskett Research and Education Area (BREA) were remeasured in 2004/2005 to assess trends in canopy composition, vegetation–environment relationships, and regeneration patterns. Quercus spp. [particularly Quercus alba (white oak)] remained dominant in BREA forests. However, Nonmetric Multidimensional Scaling Ordination of a dataset composed of tree importance values from both samples indicated the two most common community types (Dry Ridge and Slope; Mesic Slope) were shifted away from Quercus spp. abundance toward greater importance of Acer saccharum (sugar maple) in ordination space. Plots in glade-like environments shifted toward greater Juniperus virginiana (eastern redcedar) importance. Environmental analysis indicated that soil properties (pH, organic matter, Ca, Mg, and K) and slope position were highly correlated with ordination axes. Saplings of Quercus spp. exhibited greatly reduced abundance in the most recent sample and the reduction in Quercus spp. density was highest in plots with greatest canopy cover. While Acer saccharum saplings were still quite abundant in BREA forests, smaller sapling size classes were becoming notably depleted, reflecting growth of a cohort of plants established before the 1968 sample toward and into tree-sized individuals. Seedlings surveys emphasized that A. saccharum seedlings have become very rare in BREA forests, while Quercus spp. seedling densities are higher, but likely below those needed for eventual adequate recruitment to the canopy. Although a number of mechanisms to explain apparent A. saccharum regeneration failure can be developed, herbivory, recent climate trends (warming late-winter conditions) and increased litter layer depth are the most plausible explanations for the observed scarcity of A. saccharum seedlings. Although increasing importance of A. saccharum in the canopy of BREA forests appears inevitable in coming decades, longer-term trends are unclear given the shortage of seedling regeneration.

The Problem of Eastern White Pine (Pinus strobus L.) in Southern New England: Ecophysiology, Site Restriction, and Historical Land-Use Change

One of the most familiar and widespread forest tree species of eastern North America, eastern white pine (Pinus strobus L.), is also one of the least well understood. Its current ecology and distribution in southern New England comprise two very different components: even-age pure stands of old-field origin on former agricultural land, but also scattered individuals and small groups occurring in mixed stands on various classes of sites ranging from forested wetlands to dry ridge tops. A review of the historical and ecological literature of the species suggests that white pine in the presettlement forests of southern New England may have been limited to sites of shallow soil depth and low intensity of disturbance.

A 70-Year History of Arborescent Vegetation of Inwood Park, Manhattan, New York, U.S.

The arborescent vegetation located at three sites within Inwood Park, Manhattan, New York, U.S. was sampled by the quadrat method in October 2004 and May 2005 and compared with the trees present in the same quadrats on a map of trees at Inwood Park prepared by the federal Works Program Administration in 1935. Tulip poplar (Liriodendron tulipifera) was the dominant tree at the Moist Valley and South Slope sites in 1935 and 2005, whereas oak (Quercus) was the dominant genus at Dry Ridge in 1935 and 2005. Dogwood (Cornus florida) was the dominant subcanopy tree in 1935; it was unimportant in 2005, a victim of dogwood anthracnose. In terms of ecologic dominance, there has been no change in the first ranked genera at these sites in the past 70 years. Mean tree diameter (diameter at breast height) has increased from 32.3 cm (12.9 in) to 41.8 cm (16.7 in). The three sites have experienced a parallel pattern of increase in tree size from 1935 to 2005. Nonnative trees were not important in Inwood Park in 2005.

Role of evergreen understory shrub layer in the forests of the central Appalachian Highlands

Abstract Previous work indicates that an evergreen shrub layer dominated by Rhododendron maximum L. and/or Kalmia latifolia L. is a critical component of Appalachian forests, potentially exerting control over forest structure and succession. Moreover, communities characterized by thickets of evergreen understory shrubs exhibit distinct patterning on the landscape associated with topographic gradients and disturbance. We compare distributions of R. maximum and K. latifolia in the warmer and drier Ridge and Valley physiographic province and on the cooler and wetter Allegheny Plateau of western Maryland and adjacent Pennsylvania and find that the relationships between community composition and the environment differ between these two physiographic provinces. Kalmia latifolia dominance is strongly associated with topographically dry locations and a history of gypsy moth defoliation, while R. maximum is associated with topographically wet positions in the Allegheny Plateau, but less so in the Ridge and Valley....

The declining quality of late-Holocene ombrotrophic communities and the loss of Sphagnum austinii (Sull. ex Aust.) on raised bogs in Wales

The modern floristic composition of the principal raised bogs of Wales is significantly altered compared with their histories of development from the early to mid Holocene. This paper uses pollen and plant macrofossil records from two raised bogs together with previously published data from a further two sites to explore the key factors contributing to the present condition of Welsh ombrotrophic plant communities. Results show Betula and Molinia invasion is a recent feature of bog development in Wales. Previous Holocene `dry-phases' were characterized by Ericaceae and Pinus establishment, supporting recent experimental evidence that Betula and Molinia encroachment is primarily a response to nitrogen (N) loading rather than desiccation on intact bogs. Several of the bogs featured in this study also show severe signs of structural damage, not only from peat cutting but, in the case of Cors Caron, from channel processes in the adjacent River Teifi. Radiocarbon dating and inter-site comparisons of peat accumulation rates show that Rhos Goch Common in particular has been heavily impacted by peripheral peat cutting, leading to dewatering, significant subsidence of its peat dome and the spread of hummock and high dry ridge communities. Finally, the decline and local extinction of Sphagnum austinii in bogs across Europe represents one of the most significant changes in ombrotrophic community composition in the late Holocene. Co-registered pollen and macrofossil evidence from Cors Caron demonstrates that increases in landuse intensity over the last 2000 years are temporally associated with reductions in the abundance of S. austinii and ultimately its local disappearance from the palaeoecological record.

Influence of landslides on biophysical diversity — A perspective from British Columbia

Landslides have long been overlooked or underestimated as important natural disturbance agents. In particular the ecological role of landslides in maintaining biological diversity has been largely ignored. Here we provide a western Canadian ( British Columbian) perspective on the influences of landslides on biophysical diversity, which is related in several ways to biological diversity. We recognize several types of biophysical/ecological diversity: site diversity, soil diversity, and the derivative habitat or ecosystem (including aquatic ecosystems) diversity. There are also a variety of landslide types, depending on materials and on the rate and style of movement. We discuss the roles of different landslide types on various aspects of terrestrial diversity. Landslides are simultaneously depositional and erosional processes that influence sites by redistributing materials and changing surface expression — usually creating a complex microtopography that can include very dry ridges and hummocks, and sometimes depressions with standing water. Landslide impacts to site also influence soil and soil development. Portions of landslides with exposed parent material are set back to the initial stages of soil development and ecological succession. Landslides can also change soil density, structure, porosity, surface texture, chemistry and microclimate. By changing site and soil, landslides also influence habitat. Landslides influence habitat diversity by engendering a mosaic of seral stages (often both primary and secondary), and in overwhelmingly forested landscapes often create nodes or hotspots of non-forested habitat and biota. In some areas, like the boreal forest, there is an important interplay between landslides and fire, while on the coast of British Columbia debris and snow avalanches can be the dominant disturbance agent. Low-gradient and deep-seated landslides are often opportunistically colonized by beaver and other water and shrub-loving fauna. Sag ponds and impounded streams provide aquatic habitat — often with standing dead trees. Landslide rubble and scarps provide denning/nesting habitat, escape terrain, and cliff habitat for vertebrates.

Factors influencing soil invertebrate communities in riparian grasslands of the central Platte River floodplain

In the Platte River Valley of central Nebraska, USA, riparian grasslands (also known as wet meadows) have been severely impacted by a reduction in river flows, causing lower ground-water levels and altered seasonal hydroperiods. The potential impacts of these hydrologic changes, as well as the environmental factors that influence wet meadow soil invertebrate communities, are not well understood. An understanding of the ecological processes that influence these invertebrate communities is crucial for maintaining and restoring wet meadows along the Platte River. Our objectives were to describe the soil invertebrate community of wet meadows throughout the growing season and to examine the relative roles of abiotic factors in determining patterns in invertebrate community structure. We conducted the study in 12 wet meadows along the Platte River during 1999 and 2000. We identified 73 invertebrate taxa; 39 were considered soil inhabitants. Total biomass was primarily composed of earthworms, Scarabaeidae, Isopoda, and Elateridae, with earthworms and Scarabaeidae accounting for > 82%. Differences in river flow and precipitation patterns influenced some soil invertebrates. Earthworms and Scarabaeidae declined dramatically from 1999 (wet year) to 2000 (dry year). The topographic gradient created by the ridge-swale complex affected several soil invertebrate taxa; Scarabaeidae, Diplopoda, and Lepidoptera biomasses were greatest on drier ridges, while Tipulidae and Isopoda biomasses were greatest in wetter sloughs. Responses of earthworm taxa to the topographic gradient were variable, but generally, greater biomasses occurred on ridges and mid-elevations. Water-table depth and soil moisture were the most important variables influencing wet meadow soil invertebrates. Because these communities are linked to the hydrologic processes of the Platte River, future alterations of wet meadow hydrology could shift the distribution patterns of many of these invertebrates and possibly eliminate more moisture-tolerant taxa. To maintain wet meadows and their biotic communities, flow management should focus on regaining as much as possible of the former hydrograph through properly timed flows that provide an adequate hydrologie regime for wet meadows. In addition, restoration of wet meadows will depend on restoring the natural topography of wet meadows.

Leaf flushing during the dry season: the paradox of Asian monsoon forests

ABSTRACTAim Most deciduous species of dry monsoon forests in Thailand and India form new leaves 1–2 months before the first monsoon rains, during the hottest and driest part of the year around the spring equinox. Here we identify the proximate causes of this characteristic and counterintuitive ‘spring‐flushing’ of monsoon forest trees.Location Trees of 20 species were observed in semi‐deciduous dry monsoon forests of northern Thailand with a 5–6‐month‐long severe dry season and annual rainfall of 800–1500 mm. They were growing on dry ridges (dipterocarp–oak forest) or in moist gullies (mixed deciduous–evergreen forest) at 680–750 m altitude near Chiang Mai and in a dry lowland stand of Shorea siamensis in Uthai Thani province.Methods Two novel methods were developed to analyse temporal and spatial variation in vegetative dry‐season phenology indicative of differences in root access to subsoil water reserves.Results Evergreen and leaf exchanging species at cool, moist sites leafed soon after partial leaf shedding in January–February. Drought‐resistant dipterocarp species were evergreen at moist sites, deciduous at dry sites, and trees leafed soon after leaf shedding whenever subsoil water was available. Synchronous spring flushing of deciduous species around the spring equinox, as induced by increasing daylength, was common in Thailand's dipterocarp–oak forest and appears to be prevalent in Indian dry monsoon forests of the Deccan peninsula with its deep, water‐storing soils.Main conclusions In all observed species leafing during the dry season relied on subsoil water reserves, which buffer trees against prolonged climatic drought. Implicitly, rainfall periodicity, i.e. climate, is not the principal determinant of vegetative tree phenology. The establishment of new foliage before the summer rains is likely to optimize photosynthetic gain in dry monsoon forests with a relatively short, wet growing season.