Dinocyst Zones Research Articles

Late Pleistocene–Holocene palynology and paleoceanography of İmralı Basin, Marmara Sea: Pollen-spore, dinoflagellate cyst and other NPP zonations

The Marmara Sea is the oceanographic gateway that links the hypersaline Eastern Mediterranean Sea with the hyposaline Black Sea via Straits of Dardanelles and Bosphorus, respectively, and also occupies a transitional vegetation and climatic regime between the Mediterranean and Euxinic phytogeographic ecosystems. Recent palynological studies of long cores from the shallow İmralı Basin perched on the southeastern slope of the Marmara Sea reveal that major paleoecological changes accompanied the isolation and reconnection of the Marmara Sea during the last two Pleistocene glacial–interglacial cycles. Here, we use data primarily from two İmralı Basin cores to determine new quantitative zonations for the pollen-spore, dinoflagellate cyst (dinocyst) and other non-pollen palynomorph (NPP) assemblages in sediments representing the last ∼30 cal ka. A gap in one core is closed using data from three short gravity cores. Twenty-four radiocarbon ages and 20 correlation-ties for sapropel and ash layers from these five cores provide an essentially complete record for the past 30 cal ka. Use of a full suite of palynomorph data (pollen and spores, fungal spores, dinocysts and other phytoplankton, aquatic parasites/saprophytes, zoo-planktonic and zoo-benthic NPPs) shows for the first time that fluctuations in aquatic palynoassemblages during the glacial periods are more complex than simple salinity or temperature responses. The changes involve food web-level turnovers in trait- and size-based food web components in which potential parasites/saprophytes appear critically important.Pollen stratigraphies reveal a significant loss of Euxinic tree taxa by MIS 3c, with only partial recovery in MIS 1. The Euxinic forest elements are replaced by cultivars and other indicators of anthropogenic influences during the MIS 1 that is also marked by a dry interval and corresponding in part with the 4.2 cal ka Eastern Mediterranean Drought Event. The dinocyst zones for the last ∼30 cal ka show a clear turnover from assemblages dominated by brackish Ponto–Caspian taxa to marine assemblages at ∼11.7 cal ka. This phytoplanktonic signal lags slightly behind (a) the vegetation and shift in the other NPP commencing ∼13.2 cal ka, the initial marine incursion at 13.8 cal ka, and the arrival of coccolithophores at ∼13 cal ka. Comparison to published MIS 6 and 5 data reveals higher MIS 2–1 species diversity, reappearance of the MIS 6 marker Impagidinium inaequalis, and the emergences of Pyxidinopsis psilata f. quadrata, cf. Boreadinium breve, and Peridinium spp. of Brenner 2005. The pre-reconnection microbiota is dominated by the unicellular alga Sigmopollis, spikes of colonial Pediastrum algae, the saprophytes/parasites cf. Multiplicisphaeridium and Chytridiomycota. This microbiota is displaced by the post-reconnection assemblage dominated by Botryococcus coenobia, various Platyhelminth egg capsules and ostracod lining-types that persist through the Holocene. A distinctive transitional microbiota with Discocotyle-type turbellarian egg capsules marks the Bølling–Allerød warming, followed at ∼13.3 ka by a marine assemblage with micro-foraminiferal linings and crustacean eggs. The interglacial marine phase of MIS 1 in İmralı Basin has much higher dinocyst species diversity than MIS 5e, reflecting Neolithic impacts (agriculture, land clearance) and expanded shipping trade. Initial marine flooding in the Marmara Sea during both interglacials are marked by sapropel deposition but variations of ravinement surface erosion, and marine deep water residence time in a stratified Marmara Sea are potentially important factors underlying hypoxia/anoxia and sapropel deposition.

Biostratigraphy of Bajocian–Oxfordian strata based on dinoflagellate cysts and ammonites from the Dalichai Formation, northeastern Iran

The Bajocian–Oxfordian Dalichai Formation of the Binalud Mountains (northeastern Iran) consists of marls, marly limestones and limestones that were sampled and studied in detail for their palynomorph and ammonite contents. Palynological studies enabled differentiation of three marine biozones (Cribroperidinium crispum, Dichadogonyaulax sellwoodii, and Ctenidodinium continuum) within the lower interval of the formation. The palynofacies analysis shows a gradual sea-level rise during the depositional period of the rock unit, while terrestrial palynomorphs (spores and pollen grains) revealed a predominantly tropical climate for the time interval. Furthermore, a large number of ammonites were recovered from the formation, based on which 12 families, 22 genera, 30 species and 12 biozones (Zigzag, Aurigerus, Retrocostatum, Discus, Bullatus, Gracilis, Anceps, Coronatum, Athleta, Transversarium, Bifurcatus, and Bimammatum) were differentiated. When comparing the dinoflagellate cyst zones with the ammonite zones, the Dichadogonyaulax sellwoodii dinoflagellate cyst zone was correlated with the ammonite zones of Zigzag, Aurigerus, Retrocostatum, Discus, Bullatus, and Gracilis and the Ctenidodinium continuum dinocyst zone was correlated with the Gracilis and Anceps ammonite zones. The dinocyst and ammonite assemblages of the Binalud Mountains are similar to assemblages described from northern and northeastern Iran (Alborz and Koppeh-Dagh basins) as well as Northwest Europe. This suggests marine connections between these sedimentary basins during the Bajocian–Oxfordian. Consequently, the integrated dinocyst and ammonite zonation established here can be utilized also in other parts of the northern Tethyan Realm. The diachroneity of the upper boundary of the Dichadogonyaulax sellwoodii Zone between the Binalud Mountains and Northwest Europe is however revealed by comparison with the ammonite zones.

Late maastrichtian- early eocene dinoflagellate cyst biostratigraphy and aleoenvironmental study of sediments in Akukwa-1 well in the Anambra Basin

Dinoflagellate cyst biostratigraphy and palaeoenvironment of deposition of sediments penetrated by Akukwa-1 Well was carried out with a total of 253ditch cuttings using the conventional maceration technique for recovering of acid insoluble organic-walled microfossils from sediments. Sedimentological description delineated five (5) lithological units within the well, which included those of Nkporo Shale, Mamu Formation, Ajali Sandstone, Nsukka Formation and Imo Shale. A total of 14 genera and 35 species of dinoflagellate cyst were identified. The recovered dinocysts were used to establish palaeoenvironment which ranged from marginal to shallow marine, the forms were also used to propose four (4) informal dinoflagellate cyst zonations labelled (A-D) based on the first and last occurrences of two or more species. They included Zone A (Apectodinium paniculatum zone), occurring at 7990ft(Maastrichtian), zone B(Paleocystodinium australinium zone) occurring between 7990ft -4000ft (Upper Maastrichtian), zone C(Apectodinium homomorphum zone) occurring between 4000ft -2550ft (Upper Maastrichtian), and zone D(Homotryblum palladium zone) occurring between 5550ft-1520ft(Paleocene- Early Eocene). The erected dinocyst zones were correlated with the existing biozones in tropical areas of Africa and northern South America.

Dinoflagellate cyst biostratigraphy of the Bathonian, Callovian, and the Upper Jurassic in North Bulgaria

Forty-four selected organic-walled dinoflagellate cyst species have been used for biozonation of the Bathonian, Callovian, and the Upper Jurassic in North Bulgaria. The dinocysts were collected from 49 borehole sections and two exposures. The following dinocyst zones are herein introduced and described: Lithodinium valensii–Gongylodinium erymnoteichos Concurrent-range Zone (lower–middle Bathonian); Ellipsoidictyon reticulatum Assemblage Zone (upper Bathonian–lower Callovian); Dingodinium harsveldtii Taxon-range Zone (middle–upper Callovian); Lambodinium absidatum Assemblage Zone (lower Oxfordian); Scriniodinium dictyotum papillatum–Caddasphaera halosa Concurrent-range Zone (middle–upper Oxfordian); Rhyncodiniopsis gongylos Assemblage Zone (lower Kimmeridgian); Systematophora varispinosa–Gonyaulacysta? crassicornuta Concurrent-range Zone (middle–upper Kimmeridgian); Edmontodinium polyplacophorum Assemblage Zone (lower Tithonian); and Leptodinium? plagatum–Prolixosphaeridium perforospineum Concurrent-range Zone (middle–upper Tithonian). Some of the zones were partly or completely subdivided in subzones. The dinocyst zones and subzones were correlated with the Bulgarian ammonite and calpionellid zones and subzones.

Early Eocene Diatoms and Silicoflagellates of the Cis-Donets Monocline (Russian Plate): Biostratigraphical and Paleogeographical Approaches

The diatom assemblage of the Veshenskaya Formation studied in the borehole P-321 of the Cis-Donets monocline (southern part of the Russian Plate) is assigned to the lower Ypresian Moisseevia uralensis Zone. According to the available data, this zone corresponds to the Deflandrea oebisfeldensis and Dracodinium (=Wetzeliella) astra dinocyst zones and, accordingly, to the nanoplankton NP10 Zone. The presence of the zonal silicoflagellate species Naviculopsis foliacea confirms the Early Eocene age of the Veshenskaya Formation. The taxonomic similarity of diatoms and silicoflagellates of the region under consideration with silicofossil assemblages of Northern Europe suggests a stable link between these paleobasins at the beginning of the Early Eocene.

Новое местонахождение хрящевых рыб из верхнего палеоцена Донского правобережья (Волгоградская область)

An assemblage of chondrichthyans is described from the new locality near Serafimovich town (the western part of Volgograd region). More than 2500 fossils were collected from the basal gravel-pebble bed and phosphorite-bearing layer from the lower part of the Buzinovka formation (upper Paleocene). There are reworked Cretaceous and Paleocene chondrichthyans (6 species) and synchronous 21 species of sharks and 2 species of chimaeroids. The presence of synchronous Gluekmanotodus heinzelini, Odontaspis winkleri, Paraorthacodus eocaenus, Xampylodon loozi allows correlating the described lower part of the Buzinovka formation with NP8 – NP9 nannoplankton zones or Alisocysta margarita – Axiodinium augustum dinocyst zones (Thanetian – basal Ypresian).

Middle Jurassic–Lower Cretaceous stratigraphy of the northern Great Australian Superbasin: insights from maximum depositional age constraints from the U–Pb detrital zircon record

The Great Australian Superbasin, incorporating the Great Artesian Basin, is host to one of the largest artesian groundwater systems on Earth. However, despite the crucial nature of this resource to Australian agriculture, industry and rural communities, key aquifer units remain poorly studied, apart from the exposed eastern/southeastern portions of the superbasin. In this study, maximum depositional age (MDA) estimations for stratigraphic units, derived from detrital zircon U–Pb geochronology, were undertaken across the Carpentaria and northern Eromanga basins, representing the northern Queensland sector of the Great Australian Superbasin. The objective was to better constrain the ages of key aquifer and aquitard units, thereby facilitating improved stratigraphic correlations and future hydrogeological modelling. Age dating of detrital zircon significantly enhances the confidence, accuracy and precision of age assignments based solely on palynology, especially in continental facies. Furthermore, the provision of MDA estimates permits confident correlation of endemic Australian spore-pollen zonations (used to date continental strata), and dinocyst zonations (used to date marine strata) to the international time-scale. Results indicate that calculated MDAs are commonly either equivalent to, or appreciably younger than, stratigraphic age estimates based on palynostratigraphy. Therefore, several of the most economically strategic aquifer units in the superbasin have been revised upwards in geological time. Similar geochronological campaigns are required in the central, western and southern superbasin to facilitate both high-resolution correlation of key aquifer units across the superbasin and the detailed tracking of the Jurassic to Early Cretaceous paleogeographic evolution of eastern Australia. Key points U–Pb detrital zircon maximum depositional age (MDA) constraints generated are applied to the Jurassic–Cretaceous stratigraphy of the northern Great Australian Superbasin. MDA estimates facilitate robust correlation of endemic Australian palynostratigraphic zones to the international time-scale, reducing chronostratigraphic uncertainty. Several of the key aquifer units across the northern superbasin (e.g. Hooray Sandstone) yield MDA estimates younger than age assignments based on palynostratigraphy and have been revised upwards in geological time. Several units yield divergent MDA estimates, suggesting either unit diachroneity or miscorrelation, indicating that the chronostratigraphic framework of the northern superbasin (and the superbasin at large) warrants further examination.

COMPLEX BIOSTRATIGRAPHIC SCHEME OF THE JURASSIC DEPOSITS OF THE PIENNINE ZONE OF THE UKRAINIAN CARPATHIANS

The article summarizes the results of the study of paleontology and biostratigraphy of the Jurassic of the Piennine Zone of the Ukrainian Carpathians by domestic and foreign researchers. For the first time, a summary paleontological description of these sediments is presented, taking into account numerous published data on identified macro- and mictofossils. A correlation biozonal scale has been created, as well as complex biostratigraphic schemes for each Jurassic series of its region in accordance with the International Stratigraphic Scale (2020). Because the stratigraphic sequence was disturbed by disjunctive dislocations, these schemes were developed by analysis of fragmentary sections. At present, three series of the Jurassic are determined by ammonites, belemnites, pelecypods, brachiopods, the Lower and Upper series are also dated by dinocysts, the Upper also by foraminifera and tintinnids. The stages and standard zones of the modern international scale are determined by ammonites: Liasicus of the Gettangian; Bucklandi, Obtusum, Raricostatum of the Sinemurian; Jamesoni, Emaciatum of the Pliensbachian; Tenuicostatum, Serpentinum, Bifrons, Thouarcense of the Toarcien; Opalinum, Murchisonae, Bradfordensis, Concavum of the Aalenian; Propinquans, Humpriesianum, Parkinsoni of the Bajocian; Zigzag, Aurigerus, Subcontractus, Bremeri, Retrocostatum of the Bathonian; Athletа of the Callovian; Plicatilis і Bifurcates of the Oxfordian; Divisum, Acanticum і Cavouri of the Kimmeridgian. Continuous dinocyst zonation is traced in the Upper Jurassic – Fibrata Аcme (Upper Oxfordian), Parvula Acme, Moluccana і Borzai (Kimmerigian), Pulla, Tithonica, Malmica, Semiradiata (Lower Tithonian) and all standard tintinnide zones of Tithonian — Chitinoidella (підзони Dobeni і Boneti), Praetintinnopsella, Crassicollaria. The boundaries of the stages from the Gettangiean to the Oxfordian are not determined. The boundaries between the Lower and Upper Oxfordian as well as between the Kimmeridgian substages are determined by ammonites as well as the boundary between the Kimmeridgian and Tithonian is determined by dinocysts, and the boundary between the Jurassic and Cretaceous is determined by tintinnids (corresponds to the boundary between the zones Crassicollaria of Tithonian and Calpionella of Berriassian). Stratigraphic unconformities were revealed by the macro- and mictofossils: the regional erosion on the boundary between the Aalenian and Bajocian as well as in Early Callovian, and short gap on the boundary between the Lower and Upper Bajocian.

Palinostratigraphy and Paleoenvironments in the Gorodishchi Section (Middle Volga River Region, Kimmeridgian–Hauterivian)

Abstract —Eight dinocyst-based and three spore- and pollen-based biostratigraphic units are defined in the Kimmeridgian, Volgian, and Hauterivian of the Gorodishchi section, based on a biostratigraphic analysis of the successions of marine and terrestrial palynomorphs. Algological assemblages are described in more detail, and additional criteria for the definition of dinocyst zones established by previous researches are given. A more detailed biostratigraphic subdivision of the middle part of the Volgian is proposed. A local dinocyst zone in the Hauterivian and a biostratigraphic succession of spore–pollen units in the entire section are described for the first time. The research results demonstrate that the boundaries of many palynostratigraphic units exhibit a considerable correlation potential. Based on a biofacies analysis of the microphytoplankton, the dynamics of transgressive–regressive events is studied in relation to the accompanying oxygen and trophic conditions. Possible relationships between marine paleoenvironments and climatic changes reconstructed on the basis of spore–pol­len data are discussed.

Dinoflagellate cyst biostratigraphy of Upper Cretaceous turbiditic deposits from a part of the Bąkowiec section in the Skole Nappe (Outer Carpathians, southern Poland)

Dinoflagellate cysts from turbiditic deposits of the Ropianka Formation in an interval of the Bąkowiec section (Skole Nappe, Outer Carpathians, southern Poland) have been analyzed. This interval, previously attributed to the Cretaceous–Paleogene boundary transition based on planktic and benthic foraminifera, contains organic-walled dinoflagellate cysts (dinocysts) of a latest Campanian age. The difference in age, revealed here between foraminifera and dinocyst findings, is considered as an unresolved dilemma, which is pending some explanations, probably on the basis of further multidisciplinary studies envisaged in the same outcrop. The palynological content is well preserved and rich in dinocyst cyst taxa, including 37 upper Campanian index species among 92 species in total. The dinocyst assemblages observed in this interval are similar to those previously recognized in biostratigraphically well-calibrated sections of Western Europe. The dinocyst marker events enabled a correlation of the studied interval with the Samlandia mayi dinocyst Zone of Slimani (2001a), which is an equivalent to the Belemnitella minor Zone in the northwestern Europe. These dinocyst events include the first occurrences (FOs) of Kleithriasphaeridium truncatum, Nematosphaeropsi philippotii, Samlandia carnarvonensis, Samlandia mayi and last occurrences (LOs) of Corradinisphaeridium horridum, Raetiadinium truncigerum, Odontochitina operculata and Xenascus ceratioides subsp. ceratioides. The palynological content shows significant reworking of Campanian dinocyst taxa and a richness in palynodebris of continental origin, which are likely related to deposition by turbiditic currents.

Uppermost Berriasian macro- and microfossil assemblages from the Ticha Formation of the Eastern Fore-Balkan Mts (Bulgaria)

Late Berriasian macro- and microfossil assemblages from the Ticha Formation (Eastern Fore-Balkan Mts, Bulgaria) were studied. Eight ammonite taxa, including coiled and straight forms, were described. In addition, 23 calpionellid, 12 calcareous dinocyst and 4 foraminiferal taxa were identified. The “Thurmanniceras” otopeta ammonite Subzone of the Tirnovella alpillensis Zone, the calpionellid Oblonga Subzone of the Calpionellopsis Zone and the Stomiosphaera wanneri calcareous dinocyst Zone were indicated. The advent of Bochianites neocomiensis, the late occurrence of Bochianites (?) ambiguus and the earliest appearance of Phylloceras (Hypophylloceras) tethys tethys was documented in the Eastern Fore-Balkan Mts area. The calpionellid-spicule microfacies was also recognized. The fossil data allow a correlation with similar coeval localities of the European Tethyan Realm. Pelagic macro- and microfossils and the microfacies are indicative of deeper-water basin depositional settings.

A new stratigraphic framework for the Miocene – Lower Pliocene deposits offshore Scandinavia: A multiscale approach

Here, we present an updated stratigraphic subdivision of the Oligocene to Pleistocene succession (880–610 m) in the newly proposed type well for the Molo Formation, the industrial 6407/9‐5 well, located on the continental shelf in the eastern Norwegian Sea. Furthermore, new data from the Danish North Sea wells Nora‐1, Vagn‐2, and Tove‐1 are presented. The studied succession in the 6407/9‐5 well is composed of five sedimentary units separated by hiati. The dating of these five units is based on correlation to the stratigraphically more complete Neogene succession in the (Danish) central North Sea area. In this study, a robust stratigraphic framework of these five units, based on a combination of dinoflagellate cysts (dinocyst) stratigraphy and seismic data, is established. The Oligocene succession is referred to the NSO zonation of Van Simaeys et al., Review of Palaeobotany and Palynology, 2005, 134, 105–128, while the Miocene–Pliocene succession is referred to the dinocyst zonation of Dybkjær and Piasecki, Review of Palaeobotany and Palynology, 2010, 161, 1–29. In well 6407/9‐5 the two lowermost units, located below the Molo Formation, comprise a Rupelian (Lower Oligocene) succession up to 803 m, and an Aquitanian/Burdigalian (Lower Miocene) succession from 803 to 787 m, respectively. Both of these units are referred to the Brygge Formation. The Molo Formation is separated from the underlying Brygge Formation by an unconformity. Furthermore, and in contrast to previous studies, our study shows that the Molo Formation (787–703 m) has an unconformity within it. The lower part of the formation (787–724 m) is dated to late Tortonian (Late Miocene), and referred to the Hystrichosphaeropsis obscura dinocyst Zone. The upper part (724–703 m) is dated to Zanclean (Early Pliocene), and referred to the Melitasphaeridium choanophorum dinocyst Zone. The uppermost unit studied (703–670 m) in the well is referred to the Gelasian (Lower Pleistocene) and is included in the Naust Formation. The regional correlation of this stratigraphy with the complete succession in the Danish North Sea reveals that the hiati found in the Miocene succession on the Norwegian Sea shelf are controlled by tectonism, while the internal depositional patterns of the Molo Formation were controlled by eustatic sea‐level changes.

Calcareous nannofossil assemblages from Paleocene sediments of the Ukrainian Carpathians

Problem statement. Detail study of both vertical and spatial distribution of calcareous nannofossils from the Paleocene sediments of the Ukrainian Carpathians enabled to determine the evolutional development of Haptophyta algae during the Paleocene, and became the basis for correlation of boundary markers and correlative biohorizons. Here we summarize data obtained from previous research, and also our own, and propose an updad biozonation of Paleocene sediments of the Northern and Southern slopes of the Ukrainian Carpathians. Prerious work. Calcareous nannofossil research of the area goes back to the 1960’s. These studies had a fragmentary character with only few sections examined for their nannofossil content. Materials and research methods. The paper includes the results of nannofossil research from numerous sections of Paleocene Flysch developed in the Outer and Inner Carpathian nappes. Discussion of the research data. The paper discusses the biozonation and correlation of local lithostratigraphic units of the Ukrainian Carpathians based on the detail study of nannofossils. For the first time, the complete successions of nannozones of the latest Geological Time Scale have been identified in the Ukrainian Carpathians. In the Outer (Flysch) Carpathians several nannozones were determined within the Skyba and Boryslav-Pokuttya nappes. There include – Markalius inversus / Biantholithus sparsus (NP1), Cruciplacolithus tenuis (NP2), Chiasmolithus danicus (NP3) and Ellipsolithus macellus (lower NP4) in the Upper Striy subsuite; top Ellipsolithus macellus (NP4), Fasciculithus tympaniformis (NP5), Heliolithus kleinpelliі (NP6) in Yaremcha Formation, Discoaster mohleri (NP7) in Bytkiv layers, Heliolithus riedelii (NP8), Discoaster multiradiatus (bottom NP9) from the Yamna suite; Chiasmolithus danicus (NP3) from Upper Bereznyan subsuite of Dukla nappe; Fasciculithus tympaniformis (NP5) in Gnylets suite of Chornohora nappe and Markalius inversus (NP1) in Urdyn suite of Svydovets nappe. Within the Inner Carpathians nannozones were defined in the Lower Metova subsuite of the Vezhany nappe – Biantholithus sparsus (NP1), Fasciculithus tympaniformis (NP5), Heliolithus kleinpelliі (NP6), Heliolithus riedelii (NP8); Sushmanets suite of Monastyrets nappe contained Fasciculithus tympaniformis (NP5), Heliolithus kleinpelliі (NP6) and Tribrachiatus contortus (NP10) of the Early Ypresian. The stratigraphic positioning of nannofossil assemblages was rectified and their correlation with foraminifera and dinocyst zones was demonstrated. These nannozone assemblages from the Ukrainian Carpathians are correlated with the Danian, Selandian and Thanetian stages of the Geological Time Scale (GTS, 2012 and 2016). This integrated research enables to rectify and modify the stratigraphic scheme of the Paleogene from the Carpathian region. Conclusions. A modern calcareous nannofossil biozonation of the latest International schemes has been recognized from the studied Paleocene sections of the Ukrainian Carpathians. Nine (9) nannozones ranging from NP1 (Early Danian) to NP9 (Late Thanetian – Early Ypresian) and also the lowest NP10 nannozone of Early Ypresian have been identified in strata from different structural-tectonic units. The nannofossil biozonation is correlated with those foraminifera and dinocyst biochronology. The age and correlation for the Paleocene stratigraphic subdivisions has been justified based on the nannofossil biostratigraphy. This resulted in the upgrading of the existing stratigraphic scheme of the Paleogene of the Ukrainian Carpathians.

New Palynological Data from the Middle Eocene Deposits of the Kheu Reference Section (Kabardino-Balkaria, North Caucasus)

The results of palynological study of the middle–upper Eocene interval of the Kheu reference section are presented. The dinocyst zones such as Enneadocystaarcuata, Costacystabucina, Enneadocystapectiniformis, Rhombodiniumdraco, Reticulatosphaeraactinocoronata, and Talladinium? angulosum were recognized in the studied part of the section. Based on the first order calibrations with calcareous nannoplankton, their stratigraphic boundaries are established as well as the lowermost occurrences of stratigraphically important species are refined. New microplankton data suggest the early Lutetian age of the uppermost Cherkessk and Keresta formations; the Kuma Formation is dated by late Lutetian–earliest Priabonian, while the lowermost Belaya Glina Formation corresponds to the Priabonian. Analysis of ratios of marine and terrestrial palynomorphs in the Kheu section permits to interpret the paleoenvironments during the Lutetian–Priabonian in the studied part of the eastern Peri-Tethys.

Biostratigraphy and microfacies of the pelagic carbonate formations in the Yavorets section (Tithonian–Berriasian), Western Balkan Mts, Bulgaria

Calpionellid study has provided new evidence of early and late Tithonian age of the top of the Gintsi Formation, and of late Tithonian and early Berriasian age of the Glozhene Formation in the Yavorets section (Western Balkan Mts, Bulgaria). The calpionellid Chitinoidella, Praetintinnopsella, Crassicollaria, and Calpionella zones have been documented in successive order. Three calcareous dinocyst zones – Colomisphaera tenuis, Colomisphaera fortis and Stomiosphaerina proxima, have been determined in this lower Tithonian to lower Berriasian interval. From the base upwards, the following microfacies have been recognized: Saccocoma (Gintsi Formation, lower and upper Tithonian), Globochaete alpina, and calpionellid (Glozhene Formation, upper Tithonian and lower Berriasian). The base of the Berriasian has been traced at the explosion of the uniformly shaped spherical variety of Calpionella alpina. Evolutionary lineages of species of the genus Calpionella are discussed, as well as the vertical distributions and abundance peaks of crassicollarians. The calpionellid zones described herein are correlated with coeval zonations from the Western, Central and Eastern Tethyan domains. The regional correlation with previously studied sections of Tithonian/Berriasian pelagic carbonates in the Western Balkan Mts revealed a transition to hemipelagic deposition of the limestone-marl succession of the Salash Formation and/or sandstone accumulation during the middle to late Berriasian (Elliptica and Simplex calpionellid subzones) due to unstable conditions of the sedimentary environment. From the west to the east in the Western Balkan Unit (i.e., from the Rosomač section in eastern Serbia to the Sarbenitsa, Bov and Yavorets sections in the Iskar River Valley area), there is a trend of slight progressive deepening of the basin. This is manifested in the occurrence of redeposited shallow-carbonate-platform microfossils in the west to greater thickenesses of the Gintsi and Glozhene formations and occurrence of sandstone channel deposits in the east.

The Cretaceous–Paleogene (K/Pg) boundary in the Dababiya Borehole, southern Egypt: An organic-walled dinoflagellate cyst approach

For the first time, a dinoflagellate cyst analysis is provided for the Upper Cretaceous–lower Paleogene transition in the Dababiya borehole, near Luxor, southern Egypt. Rich and relatively well-preserved dinoflagellate cyst assemblages have been recovered; a total of 173 species and subspecies have been identified. Many of these were not previously known from Egyptian material. The biostratigraphical dinoflagellate cyst events recorded in the present study have allowed us to propose the first K/Pg boundary dinocyst zonation scheme for Egypt and to compare this with existing dinoflagellate cyst zonations worldwide. In ascending order, the observed latest Maastrichtian dinoflagellate cyst events include successively the last occurrence (LO) of Cannosphaeropsis utinensis, the first occurrences (FOs) of Apteodinium fallax, Disphaerogena carposphaeropsis, Kenleyia leptocerata and the LOs of Alisogymnium euclaense, Dinogymnium spp. and Pterodinium cretaceum. The most significant early Danian dinoflagellate cyst events used in this study are the FOs of Danea californica, Carpatella cornuta, Cassidium fragile, Lanternosphaeridium reinhardtii, Senoniasphaera inornata and Membranilarnacia? tenella. The dinoflagellate cyst biostratigraphical results, calibrated with data from benthic and planktonic foraminifera and calcareous nannoplankton from the same set of samples, have enabled a more accurate identification of the Cretaceous–Paleogene (K/Pg) boundary than previously suggested for the succession studied.

Palynological study of paleoclimate and paleoceanographic changes in the Eastern South Korea Plateau, East Sea, during the Plio-Pleistocene climate transition

A palynological study was performed using the core obtained from the Integrated Ocean Drilling Program (IODP) Expedition 346 Site U1430 in the Eastern South Korea Plateau in the East Sea, to reveal variability of paleoclimate and paleoceanography during the late Pliocene to early Pleistocene climate transition period (56.0–68.0 meters below sea floor [mbsf]). Two pollen and dinocyst zones are established with a boundary at ca. 3.0 Ma. Prior to ca. 3.0 Ma (first pollen and dinocyst zone), long-term oscillation of the East-Asian winter monsoon (EAWM) is indicated by fluctuations in the abundance of pollen taxa, representing cold–dry and cool–wet climates. After ca. 3.0 Ma (second pollen and dinocyst zone), an increase in cold–dry conifer pollen indicates an enhanced and stable EAWM. In addition, the prevalence of the xerophytic herb, Artemisia, increases markedly after 3.0 Ma, suggesting open grassland on the exposed South Yellow Sea shelf during the sea level lowstand under cold–dry climate conditions. Artemisia pollen are interpreted as having been delivered to the coast through fluvial channels in this grassland environment, and subsequently transported to the East Sea by inflow of a weak Tsushima Warm Current (TWC) through the narrow Korea Strait. The TWC intrusion signal is represented by an influx of Tuberculodinium vancampoae, a warm water dinoflagellate species, in the western Pacific region. The high abundances of cold–dry conifer pollen and cold water dinocyst elements after ca. 2.8 Ma suggest an intensified EAWM. Meanwhile, a short-term increase in moisture-loving Tsuga pollen after ca. 2.5 Ma was likely due to soil moisture derived from snowfall caused by evaporation of heated seawater from the weak TWC in the East Sea under cold climate conditions. The palynological studies reported here help verify the linkages between the EAWM, TWC, precipitation and vegetation responses in/around the East Sea region.

An updated and revised stratigraphic framework for the Miocene and earliest Pliocene strata of the Roer Valley Graben and adjacent blocks

AbstractIn the Netherlands, the bulk of the Miocene to lowest Pliocene sedimentary succession is currently assigned to a single lithostratigraphical unit, the Breda Formation. Although the formation was introduced over 40 years ago, the definition of its lower and upper boundaries is still problematic. Well-log correlations show that the improved lecto-stratotype for the Breda Formation in well Groote Heide partly overlaps with the additional reference section of the older Veldhoven Formation in the nearby well Broekhuizenvorst. The distinction between the Breda and the overlying Oosterhout Formation, which was mainly based on quantitative differences in glauconite and molluscs, gives rise to ongoing discussion, in particular due to the varying concentrations of glauconitic content that occur within both formations. In addition, the Breda Formation lacks a regional-scale stratigraphic framework which relates its various regionally to locally defined shallow marine to continental members.In order to resolve these issues, we performed renewed analyses of material from several archived cores. The results of archived and new dinocyst analyses were combined with lithological descriptions and wire-line log correlations of multiple wells, including the wells Groote Heide and Broekhuizenvorst. In this process, the updated dinocyst zonation of Munsterman & Brinkhuis (2004), recalibrated to the Geological Time Scale of Ogg et al. (2016), was used. To establish regionally consistent lithostratigraphic boundaries, additional data was used along a transect across the Roer Valley Graben running from its central part (well St-Michielsgestel-1) towards the southern rift shoulders (well Goirle-1). Along this transect, chronostratigraphic and lithostratigraphic analyses were integrated with well-log correlation and the analyses of seismic reflection data to constrain geometrical/structural relationships as well.The results led to the differentiation of two distinct seismic sequences distinguished by three recognisable unconformities: the Early Miocene Unconformity (EMU), the Mid-Miocene Unconformity (MMU) and the Late Miocene Unconformity (LMU). The major regional hiatus, referred to as the Mid-Miocene Unconformity, occurs intercalated within the present Breda Formation and compels subdivision of this unit into two formations, viz. the here newly established Groote Heide and the younger Diessen formations. Pending further studies, the former Breda Formation will be temporally raised in rank to the newly established Hilvarenbeek subgroup, which comprises both the Groote Heide and Diessen formations. Whereas these two sequences were already locally defined, a third sequence overlying the LMU represents two newly defined lithostratigraphical units, named the Goirle and the Tilburg members, positioned in this study at the base of the Oosterhout Formation. Besides their unique lithological characteristics, in seismic reflection profiles the Goirle and the Tilburg members stand out because of their distinct seismic facies.Use of an integrated, multidisciplinary and regional approach, an improved southern North Sea framework and more comprehensive lithostratigraphic subdivision of Neogene successions is proposed for the Netherlands, to make (cross-border) correlations more straightforward in the future.

Identification of the Paleocene–Eocene boundary in coastal strata in the Otway Basin, Victoria, Australia

Abstract. Detailed, stratigraphically well-constrained environmental reconstructions are available for Paleocene and Eocene strata at a range of sites in the southwest Pacific Ocean (New Zealand and East Tasman Plateau; ETP) and Integrated Ocean Discovery Program (IODP) Site U1356 in the south of the Australo-Antarctic Gulf (AAG). These reconstructions have revealed a large discrepancy between temperature proxy data and climate models in this region, suggesting a crucial error in model, proxy data or both. To resolve the origin of this discrepancy, detailed reconstructions are needed from both sides of the Tasmanian Gateway. Paleocene–Eocene sedimentary archives from the west of the Tasmanian Gateway have unfortunately remained scarce (only IODP Site U1356), and no well-dated successions are available for the northern sector of the AAG. Here we present new stratigraphic data for upper Paleocene and lower Eocene strata from the Otway Basin, southeast Australia, on the (north)west side of the Tasmanian Gateway. We analyzed sediments recovered from exploration drilling (Latrobe-1 drill core) and outcrop sampling (Point Margaret) and performed high-resolution carbon isotope geochemistry of bulk organic matter and dinoflagellate cyst (dinocyst) and pollen biostratigraphy on sediments from the regional lithostratigraphic units, including the Pebble Point Formation, Pember Mudstone and Dilwyn Formation. Pollen and dinocyst assemblages are assigned to previously established Australian pollen and dinocyst zonations and tied to available zonations for the SW Pacific. Based on our dinocyst stratigraphy and previously published planktic foraminifer biostratigraphy, the Pebble Point Formation at Point Margaret is dated to the latest Paleocene. The globally synchronous negative carbon isotope excursion that marks the Paleocene–Eocene boundary is identified within the top part of the Pember Mudstone in the Latrobe-1 borehole and at Point Margaret. However, the high abundances of the dinocyst Apectodinium prior to this negative carbon isotope excursion prohibit a direct correlation of this regional bio-event with the quasi-global Apectodinium acme at the Paleocene–Eocene Thermal Maximum (PETM; 56 Ma). Therefore, the first occurrence of the pollen species Spinizonocolpites prominatus and the dinocyst species Florentinia reichartii are here designated as regional markers for the PETM. In the Latrobe-1 drill core, dinocyst biostratigraphy further indicates that the early Eocene (∼ 56–51 Ma) sediments are truncated by a ∼ 10 Myr long hiatus overlain by middle Eocene (∼ 40 Ma) strata. These sedimentary archives from southeast Australia may prove key in resolving the model–data discrepancy in this region, and the new stratigraphic data presented here allow for detailed comparisons between paleoclimate records on both sides of the Tasmanian Gateway.

Upper Jurassic foraminifera, dinoflagellates and terrestrial sporomorphs from the Gorodishche Section (Ul’yanovsk Region, East European Platform, Russia): Biostratigraphic, palaeoenvironmental and palaeobiogeographical implications

The Upper Kimmeridgian to Lower Volgian successions of the East European Platform are well represented in the clay formations of the Gorodishche Section (Ul’yanovsk Region). It displays very rich foraminiferal and palynological assemblages. A review of foraminiferal and microphytoplankton biozonation is proposed in this study. Foraminiferal as well as marine and terrestrial palynomorph assemblages were also analyzed to reconstruct environmental and climatic conditions during the transition from the Late Kimmeridgian to the Early Volgian. The depositional setting corresponds to the transition from the upper to the middle sublittoral zone mostly associated with oxic conditions. In addition, facies analysis allows the reconstruction of short term sea-level curve. Curves of the Jaccard similarity Index have been constructed between the Gorodishche Section and different regions of Eurasia in order to understand palaeobiogeographical changes. The results of the analyses indicate significant similarities of foraminiferal and dinocysts associations across East European Platform, Western Europe, and Siberia probably associated to good connections between palaeobasins. The wide geographical range of foraminiferal and dinocyst zones indicates the possibility for long distance correlations in the latest Kimmeridgian probably associated with palaeoenvironmental or/and palaeogeographical conditions favorable for the marine biota exchange. Finally, palaeoclimatic reconstructions are derived from the analysis of spore–pollen associations. They indicate a warm arid to semi-arid climate with alternations of wet and dry seasons. The nature of the near-shore land vegetation was strongly influenced by humidity fluctuations.