Cai, C., Clarke, D.J., Huang, D. & Nel, A., 2014. A new genus and species of Steninae from the late Eocene of France (Coleoptera, Staphylinidae). Alcheringa 38, 557–562. ISSN 0311-5518.A remarkable new genus and species of rove beetle, Eocenostenus fossilis gen. et sp. nov., is described and figured based on one well-preserved specimen from the late Eocene of Monteils (near Alès, Gard, France). Eocenostenus is definitively placed in the extant subfamily Steninae, based on the combination of dense and coarse body punctation, globular and protruding eyes, exposed and closely spaced antennal insertions on the vertex, and six visible abdominal terga. Eocenostenus differs from the two extant stenine genera Stenus and Dianous most notably in the structure of the prothorax, which is strongly transverse and with unusual anterolateral projections, and in the anteriorly placed antennal insertions. This new discovery highlights the palaeodiversity of a genus-poor subfamily and suggests that the early diversification of Steninae is probably complicated.Chenyang Cai [caichenyang1988@163.com] and Diying Huang [dyhuang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Rd., Nanjing 210008, PR China; Dave J Clarke [dclarke@fieldmuseum.org], Integrative Research Center, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA; and André Nel [anel@mnhn.fr], Muséum National d’Histoire Naturelle, Institut de Systématique, Evolution, Biodiversité, ISYEB, UMR 7205 CNRS UPMC EPHE, CP50, 45 rue Buffon, F-75005 Paris, France. Received 23.1.2014; revised 2.5.2014; accepted 12.5.2014.