The 2023 discovery of a fish fossil from lower Eocene strata of the Chuckanut Formation provides new insights into the paleoenvironment and paleoecology of one of the region’s most prolific fossil deposits. The detrital clastic fluvial and floodplain deposits of the Chuckanut Formation are not favorable for the preservation of fish, but the high quality of preservation of this specimen is evidence that some Chuckanut Formation sediments provide suitable depositional conditions for the preservation of skeletal remains. This information improves our understanding of the range of depositional environments within the Chuckanut Formation, and provides clues for searching for additional specimens. The discovery of this fossil has larger significance; the skeletal remains of fish are scarce in fluvial and floodplain deposits. Despite its incompleteness, dorsal fin and caudal fin ray anatomy suggest that the specimens represent the extinct genus Phareodus, an open-water carnivore that has previously only been reported in North America from the Green River and Bridger Formations in Wyoming and Utah, USA.