Brushy Basin Research Articles

Description of a new specimen of Haplocanthosaurus from the Dry Mesa Dinosaur Quarry.

A new specimen of Haplocanthosaurus is described based on bones excavated from the Late Jurassic Dry Mesa Dinosaur Quarry near Delta, Colorado. The specimen consists of seven dorsal vertebrae and a right tibia and is identified as Haplocanthosaurus based on the dorsally angled transverse processes, tall neural arch peduncles, low parapophyses relative to the diapophyses in the posterior dorsal vertebrae, and the robustness of the tibia combined with a greatly expanded distal articular surface. The discovery adds to our understanding of the biostratigraphy of Haplocanthosaurus, showing this genus is definitively present in the Brushy Basin Member of the Morrison Formation, and making this individual the geologically youngest Haplocanthosaurus specimen on the Colorado Plateau. The identification of this genus adds to the known diversity of sauropods at Dry Mesa Dinosaur Quarry (DMDQ), which is at least six distinct genera, making DMDQ the most diverse single locality of sauropods in the Morrison Formation and the world.

Sandstone-hosted uranium deposits of the Colorado Plateau, USA

More than 4,000 sandstone-hosted uranium occurrences host over 1.2 billion pounds of mined and in situ U3O8 throughout the Colorado Plateau. Most of the resources are in two distinct mineral systems with deposits hosted in the Triassic Chinle and Jurassic Morrison Formations. In the Chinle mineral system, base metal sulfides typically accompany mineralization. The Morrison mineral system is characterized by V/U ratios up to 20. The uranium source was likely volcanic ash preserved as bentonitic mudstones in the Brushy Basin Member of the Morrison Formation, and lithic volcanic clasts, ash shards, and bentonitic clay in the lower part of the Chinle Formation. Vanadium originated from two possible sources: iron–titanium oxides that are extensively altered in bleached rock near deposits or from similar minerals in variably bleached red beds interbedded with and beneath the Morrison. In Chinle-hosted deposits, in addition to volcanic ash, a contributing source of both vanadium and uranium is proposed here for the first time to be underlying red beds in the Moenkopi and Cutler Formations that have undergone a cycle of reddening-bleaching-reoxidation. Transport in both systems was likely in groundwater through the more permeable sandstones and conglomerate units. The association of uranium minerals with carbonate and more rarely apatite, suggests that transport of uranium was as a carbonate or phosphate complex. The first comprehensive examination of paleoclimate, paleotopography, and subsurface structure of aquifers coupled with analysis of the geochronology of deposits suggests that that there were distinct pulses of uranium mineralization/redistribution during the period from about 259 Ma to 12 Ma when oxidized mineralizing fluids were intermittently rejuvenated in the Plateau in response to changes in tectonic regime and climate. Multiple lines of evidence indicate that deposits formed at ambient temperatures of about 25 °C to no greater than about 140 °C. In both systems, deposits formed where groundwater flow slowed and was subject to evaporative concentration. Stagnant conditions allowed for prolonged interaction of U- and V-enriched groundwater with ferrous iron-bearing reductants, such as illite and iron–titanium oxides, and more rarely organic material such as plant debris. Paragenetically late in the sequence, reducing fluids introduced additional organic matter to some deposits. Reducing fluids and introduced organic matter (now amorphous and altered by radiolysis) may originate from regional petroleum systems where peak oil and gas generation was from ∼ 82 to ∼ 5 Ma. Our novel analysis indicates that these reducing fluids bleached rock and protected affected deposits from remobilization during exposure and weathering that followed uplift of the Plateau (∼80 to 40 Ma).

PRESERVED MEMBRANE ON DINOSAUR EGGSHELL FRAGMENTS, UPPER JURASSIC MORRISON FORMATION, EASTERN UTAH

ABSTRACT Dinosaur eggshell fragments, from the Upper Jurassic Brushy Basin Member of the Morrison Formation, Utah, were examined using Field Emission Scanning Electron Microscope (FESEM), Energy Dispersive Spectroscopy, and Raman Spectroscopy. Analyses revealed that the mammillary tips on the shell interior contain carbonaceous residue. Comparison under the FESEM of these shells with modern bird shells, including some samples heated to diagenetic temperatures, indicate that the residue is degraded organic compounds (DOC). Bird egg membrane is composed of interlaced collagen fibers. Features observed on, and common to, modern bird and dinosaur egg fragments include: (1) irregular-shaped calcium carbonate grains “floating” in an organic matrix; (2) three-dimensional organic fiber matrix; (3) external calcium carbonate molds of fibers in the mammillary bodies; and in heated specimens, (4) carbonaceous residue with ovate to circular pores. However, unlike birds' eggs, the dinosaur eggs contain a calcium carbonate tube around fibrous organic material that emerges from the tube and spreads laterally in a ‘puddle-like' deposit. The sizes of circular organic matrix pores of the dinosaur egg fragments are significantly smaller than those in the bird shells. Gallus gallus domesticus eggshell membranes heated to diagenetic temperatures resulted in alteration of collagen fibers to gel-like substances. The organic matrix with ovate to circular pore openings and the puddle-like deposits in the dinosaur egg fragments are interpreted as the product of membrane thermal diagenesis. The recognition of carbonaceous residue of the shell membrane on dinosaur shell fragments opens newfound opportunities to explore DOC associated with fragmental dinosaur eggs.

Two-seeded cones of probable gnetalean affinity from the Morrison Formation (Late Jurassic) of Utah and Colorado, USA

Three dimensional calcitic casts of a two-seeded compound cone are described from the Upper Jurassic Brushy Basin Member of the Morrison Formation based on specimens from Colorado and Utah. Cones of Bassitheca hoodiorum gen. et sp. nov. are broadly obovate in face view, ~3.5 mm high, ~2.6 mm wide, and viewed from above have two planes of symmetry. Micro-CT scanning of numerous specimens shows that each cone has two well-developed orthotropous seeds positioned opposite each other in the major plane and separated by a median longitudinal septum in the minor plane. At a higher level in the cone, a median apical septum in the major plane separates two depressions arranged opposite each other. The two well-developed seeds in the major plane are enclosed by bracts, but numerous incomplete cones, together with cones showing lines of dehiscence, indicate that each seed was shed by the abscission of a lateral valve. One specimen indicates that three vascular bundles entered the base of each well-developed seed and these are interpreted as the vascular supply to the envelope of a chlamydospermous ovule. The opposite and decussate arrangement of bracts at the base of the cone, the paired arrangement of the two well-developed seeds perpendicular to the two apical depressions, combined with the inferred seed envelope that was apparently shed with the seed, indicates a probable relationship to Gnetales and specifically to extant Ephedra. Along with Dayvaultia tetragona Manchester et Crane, also known from the Brushy Basin Member of the Morrison Formation, Bassitheca hoodiorum provides further evidence that the Gnetales were more significant in the Morrison vegetation than has been recognized previously.

Distinctive quadrangular seed-bearing structures of gnetalean affinity from the Late Jurassic Morrison Formation of Utah, USA

A new kind of seed-bearing structure is described based on three-dimensional casts and partially permineralized small cones from the Upper Jurassic Brushy Basin Member of the Morrison Formation, in the Henry Mountains of Utah. Cones of Dayvaultia tetragona gen. et sp. nov. are obovate in lateral view, 10.0–11.0 mm long, square in cross-section and 5.1–8.0 mm wide, with a thick wall composed of four tightly adhering bracts that open apically to expose the tips of six or eight elongate, four-lobed seeds. Micro-CT scanning reveals that the seeds are borne on a cup-shaped receptacle in a regular opposite and decussate manner. This regular arrangement, as well as similarities of the seeds to several kinds of Early Cretaceous chlamydospermous seeds, including those of Lobospermum and Battenispermum, suggests a relationship to extant and Cretaceous members of Gnetales. The sedimentary context in which the cones occur, combined with their local abundance, suggests that Dayvaultia was common on intermittently inundated well-drained floodplains during Morrison times, enhancing insight into the vegetation that supported the diverse vertebrate faunas for which the Morrison Formation is well known. http://zoobank.org/urn:lsid:zoobank.org:pub:0ECD4B37-E6B6-4050-B45A-28D713321EB8

Stratigraphy and Sedimentology of the Morrison Formation in the Western Panhandle of Oklahoma with Reference to the Historical Stovall Dinosaur Quarries

New investigations of the Morrison Formation in the western Oklahoma panhandle reveal that the formation is approximately 60 m thick near Kenton, considerably less than historical measurements. We provide a new isopach map of the Morrison Formation at the basin margin and divide the formation into three new members based on geological, geochemical, petrographic, and paleontological characteristics. Each member is defined by unique depositional facies recording a variable climatic signal and the eventual progradation of the distributive fluvial system to the basin margin. Analogous depositional facies and lithologies of these units suggest they are lateral facies successions of the well-established members of the Colorado Plateau. The Cimarron Member is composed of numerous small, ephemeral, clastic lakes at the distal margin of the alluvial braid plain of the Tidwell Member. The Boise Member consists of numerous perennial limestone lakes at the distal edge of the Salt Wash Member distributive fluvial system. At the top of the member, the lake coalesced into a large lake, herein termed Lake Stovall. The Kenton Member records the progradation of the Brushy Basin Member distributive fluvial system to the basin margin in Oklahoma. Kenton Member fluvial deposition was by small, isolated, anastomosing fluvial channels and their associated splays. The gradual progradation of facies to the basin margin signifies that deposition in the foreland basin was geographically extensive and protracted. Dinosaur fossils excavated during the 1930s are placed into a stratigraphic framework. All quarries lie in the basal 6 m of the Kenton Member. Depositional facies and taphonomic data imply the dinosaurs died during a series of severe droughts. Bone dispersal and burial resulted from ephemeral flood splay events.

New social insect nests from the Upper Jurassic Morrison Formation of Utah

This paper reports a new assemblage of social insect ichnofossils from the Brushy Basin Member of the Upper Jurassic Morrison Formation near Green River, Utah. At least seven distinct nests are visible in the locality horizon, identifiable at the outcrop scale by loci of anastomosing, and orthogonally connected hor-izontal burrows and vertical shafts. A boulder-sized block from the in situ horizon has eroded and rolled downhill, revealing the ventral aspect of the nest, showing a view of the overall nest architecture. Burrow and shaft clusters are organized into mega-galleries which have branching arms and ovate, bulbous cham-bers. The organization of distinct trace morphologies is consistent with ethological complexity of the social insects. A small sample was collected and analyzed by serial sectioning and petrographic thin sectioning to observe small-scale morphological features. Centimeter-scale analysis shows chamber, gallery, and burrow walls have complex topography. Pebble-sized, hollow, ellipsoid features are distributed throughout the up-permost facies of the nest and have undergone complete silicification of their outer surfaces. The ellipsoids share similarity with pellet structures made of mud or carton produced by modern termites. This trace fossil assemblage suggests it is possible that termites had acquired subterranean nesting behavior, and mud or carton utilization in nest construction in seasonally arid habitats by the Late Jurassic.

Decomposition of dinosaurian remains inferred by invertebrate traces on vertebrate bone reveal new insights into Late Jurassic ecology, decay, and climate in western Colorado.

A survey of 2,368 vertebrate fossils from the Upper Jurassic Mygatt-Moore Quarry (MMQ) (Morrison Formation, Brushy Basin Member) in western Colorado revealed 2,161 bone surface modifications on 884 specimens. This is the largest, site-wide bone surface modification survey of any Jurassic locality. Traces made by invertebrate actors were common in the assemblage, second in observed frequency after vertebrate bite marks. Invertebrate traces are found on 16.174% of the total surveyed material and comprise 20.148% of all identified traces. Six distinct invertebrate trace types were identified, including pits and furrows, rosettes, two types of bioglyph scrapes, bore holes and chambers. A minimum of four trace makers are indicated by the types, sizes and morphologies of the traces. Potential trace makers are inferred to be dermestid or clerid beetles, gastropods, an unknown necrophagous insect, and an unknown osteophagus insect. Of these, only gastropods are preserved at the site as body fossils. The remaining potential trace makers are part of the hidden paleodiversity from the North American Late Jurassic Period, revealed only through this ichnologic and taphonomic analysis. Site taphonomy suggests variable, but generally slow burial rates that range from months up to 6 years, while invertebrate traces on exposed elements indicate a minimum residence time of five months for carcasses with even few preserved invertebrate traces. These traces provide insight into the paleoecology, paleoclimate, and site formation of the MMQ, especially with regards to residence times of the skeletal remains on the paleolandscape. Comprehensive taphonomic studies, like this survey, are useful in exploring patterns of paleoecology and site formation, but they are also rare in Mesozoic assemblages. Additional work is required to determine if 16.174% is typical of bulk-collected fossils from Jurassic ecosystems in North America, or if the MMQ represents an unusual locality.

The Morrison Formation and its bounding strata on the western side of the Blanding basin, San Juan County, Utah

In 2016 and 2017, the Utah Geological Survey partnered with the U.S. Bureau of Land Management to conduct a paleontological inventory of the Morrison Formation south and west of Blanding, Utah, along the eastern margin of the Bears Ears National Monument. The Morrison in this region is critical to understanding Upper Jurassic stratigraphy across the Colorado Plateau because it is the type area for the Bluff Sandstone, Recapture, Westwater Canyon, and Brushy Basin Members of the Morrison Formation, which are the basis for nomenclature in New Mexico and Arizona as well. Researchers have disagreed about nomenclature and correlation of these units, which transition northward in the study area into the Tidwell, Salt Wash, and Brushy Basin Members. Numerous vertebrate localities make inclusion of the Bluff Sandstone and Recapture Members in the Middle Jurassic San Rafael Group, as suggested by some previous workers, unlikely. The Salt Wash Member does not separate the Bluff Sandstone and Recapture Members at Recapture Wash, but sandstone lenses of Salt Wash facies occur higher in northern Recapture exposures. Northward, along the outcrop belt east of Comb Ridge, the Bluff-Recapture interval thins, interlenses, and pinches out into the Tidwell and lower Salt Wash, with the main lower sandstone interval of the Westwater Canyon merging northward into the upper Salt Wash Member.
 The partly covered, 1938 type section of the Brushy Basin Member is identified along Elk Mountain Road at the southern end of Brushy Basin. We describe a detailed, accessible Morrison Formation reference section about 11.2 km (7 mi) to the south along Butler Wash. There, 81.68 m (268 ft) of Brushy Basin Member is well exposed along a road between the top of the Westwater Canyon Member and the base of the Lower Cretaceous Burro Canyon Formation. We informally call the upper sandstone bed(s) of the Westwater Canyon Member that cap mesas and benches in the region “No-Mans Island beds.” Smectitic mudstones between the No-Mans Island beds and the main sandstone body of the Westwater Canyon suggest that the Salt Wash-Brushy Basin contact to the north may be somewhat older than the base of the Brushy Basin Member as originally defined in its type area. Determining whether the No-Mans Island beds pinch out to the north or are removed by erosion below the regional basal Brushy Basin paleosol requires further research. Several significant fossil vertebrate and plant sites have been documented in the Brushy Basin type area. Newly identified volcanic ashes provided zircons for U-Pb ages of 150.67 ± 0.32 Ma from near the top of the Brushy Basin Member and of 153.7 ± 2.1 Ma and 153.8 ± 2.2 Ma for two zircons in lower part of Recapture Member. At the top of the Brushy Basin Member, ferruginous paleosols commonly overlying conglomeratic sandstone are speculated to be of Early Cretaceous age (detrital zircon age pending) and are assigned herein to the Yellow Cat Member of the Burro Canyon Formation. These iron-rich paleosols suggest wetter climatic conditions during the Jurassic-Cretaceous transition in the Blanding basin.

Stratigraphic setting of fossil log sites in the Morrison Formation (Upper Jurassic) near Dinosaur National Monument, Uintah County, Utah, USA

The outcrop belt of the Upper Jurassic Morrison Formation in the northeastern Uinta Basin and southeastern flank of the Uinta Mountains is particularly rich in dinosaurian and non-dinosaurian faunas, as well as in fossil plants. The discovery of several well-preserved, relatively intact, fossil logs at several locations in Rainbow Draw and one location in Miners Draw, both near Dinosaur National Monument (Utah), has provided an opportunity to study the local paleobotany, stratigraphy, and sedimentology of the Morrison Formation in northeastern Utah. The Morrison Formation in northeastern Utah consists of four members. In ascending chronostratigraphic order, they are the Windy Hill, Tidwell, Salt Wash, and Brushy Basin Members. The lithology (including the presenceof glauconite grains) and fossil assemblage of the lower two members (Windy Hill and Tidwell) indicate a marine to marginal marine (coastal plain) depositional environment, whereas the lithology, fossil flora and faunaassemblage of the upper two members (Salt Wash and Brushy Basin) indicate a fluvial–lacustrine depositional environment. At least 10 fossil log sites in Rainbow Draw have been documented so far, and geologic mapping indicates that the logs and wood all occur in the same stratigraphic interval within the Salt Wash Member, approximately 17 to 27 m above the base of the member. The unit containing the logs and wood is about 11 m thick and consists of very fine to fine-grained sandstone and siltstone with indistinct bedding and no discernible sedimentary features.The logs are siliceous, some have a coaly exterior, and they range in exposed length from 0.5 to 11 m and reach diameters up to 1.1 m. In the Miners Draw area, a single siliceous log is documented in the upper part of the Salt Wash Member within a silty sandstone unit that is 4 m thick; its exposed length is about 6 m. Although the correlation of the Miners Draw log-bearing interval to the interval in Rainbow Draw is uncertain, both units are lithologically similar and both occur in the upper part of the Salt Wash Member. The logs have been identified as araucariaceous conifers that pertain to the same taxon originally described as Araucarioxylon hoodii Tidwell et Medlyn 1993 from Mt. Ellen in the Henry Mountains of southern Utah. Concurrent systematic work will prompt a nomenclatural transfer of this species to the genus Agathoxylon. Based on the abundance of large fossil logs and wood in the same stratigraphic interval in Rainbow Draw, wehypothesize that the area was covered by stands of moderately large trees of araucariaceous conifers. The sedimentological evidence suggests that the trees were not transported far from their original site of growth before they were deposited in a low-energy floodplain environment.

New Chemical Abrasion Thermal Ionization Mass Spectrometry Dates from the Brushy Basin Member, Morrison Formation, Western Colorado: Implications for Dinosaur Evolution

AbstractTwo new chemical abrasion thermal ionization mass spectrometry U/Pb dates establish the age of the top of the Brushy Basin Member, Morrison Formation, at Flat Top (also known as Terrace Hil...

The Upper Jurassic Morrison Formation in north-central New Mexico–Linking Colorado Plateau stratigraphy to the stratigraphy of the High Plains

Most study of the Upper Jurassic Morrison Formation has focused on its spectacular and extensive outcrops on the southern Colorado Plateau. Nevertheless, outcrops of the Morrison Formation extend far off the Colorado Plateau, onto the southern High Plains as far east as western Oklahoma. Outcrops of the Morrison Formation east of and along the eastern flank of the Rio Grande rift in north-central New Mexico (Sandoval, Bernalillo, and San­ta Fe Counties) are geographically intermediate between the Morrison Formation outcrops on the southeastern Colorado Plateau in northwestern New Mexico and on the southern High Plains of eastern New Mexico. Previous lithostratigraphic correlations between the Colorado Plateau and High Plains Morrison Formation outcrops using the north-central New Mexico sections encompassed a geographic gap in outcrop data of about 100 km. New data on previously unstudied Morrison Formation outcrops at Placitas in Sandoval County and south of Lamy in Santa Fe County reduce that gap and significantly add to stratigraphic coverage. At Placitas, the Morrison Formation is about 141 m thick, in the Lamy area it is about 232 m thick, and, at both locations, it consists of the (ascending) sandstone-dominated Salt Wash Member, mudstone-dominated Brushy Basin Member, and sandstone-dominat­ed Jackpile Member. Correlation of Morrison strata across northern New Mexico documents the continuity of the Morrison depositional systems from the Colorado Plateau eastward onto the southern High Plains. Along this transect, there is significant stratigraphic relief on the base of the Salt Wash Member (J-5 unconformity), the base of the Jackpile Member, and the base of the Cretaceous strata that overlie the Morrison Formation (K unconfor­mity). Salt Wash Member deposition was generally by easterly-flowing rivers, and this river system continued well east of the Colorado Plateau. The continuity of the Brushy Basin Member, and its characteristic zeolite-rich clay facies, onto the High Plains suggests that localized depositional models (e.g., “Lake T’oo’dichi’) need to be re-eval­uated. Instead, envisioning Brushy Basin Member deposition on a vast muddy floodplain, with some localized lacustrine and palustrine depocenters, better interprets its distribution and facies.

Paleontology, taphonomy, and sedimentology of the Mygatt-Moore Quarry, a large dinosaur bonebed in the Morrison Formation, western Colorado—Implications for Upper Jurassic dinosaur preservation modes

The Mygatt-Moore Quarry is a deposit of several thousand dinosaur bones in the Brushy Basin Member of the Morrison Formation in western Colorado. The site has been worked for more than 30 years and nearly 2400 mapped specimens have been collected. This study gathered data about the quarry from many sources to investigate the origin of the deposit. The Mygatt-Moore Quarry appears to be an attritional deposit of a relatively restricted diversity of dinosaurs, with few other non-dinosaurian taxa, that accumulated in a vernal pool deposit in an overbank setting. Bone modification was mostly by corrosion and breakage by trampling; scavenging was abundant. The paleofauna is dominated by Allosaurus and Apatosaurus (MNI and NIS), with the polacanthid ankylosaur Mymoorapelta less common. The matrix of the main quarry layer includes abundant carbonized fragments of plant material, and the mud during the time of deposition may have been often at least damp and occasionally acidic and dysoxic. The Cleveland-Lloyd Dinosaur Quarry is a close correlate of the Mygatt-Moore Quarry in terms of lithology and taphonomy, but demonstrates significant differences upon close inspection of matrix details and bone modification. Large quarries of fine-grained facies in the Morrison Formation possess a very different preservation mode as well as different taxon and relative abundance profiles from those in coarser sediments, which suggests that more may be learned in the future from taphofacies study of large quarries in mudstone beds.

Evidence for niche partitioning among ground-height browsing sauropods from the Upper Jurassic Morrison Formation of North America

Two tooth-bearing snout fragments from a diplodocid sauropod from the Brushy Basin Member of the Morrison Formation (Upper Jurassic) excavated from the Mygatt-Moore Quarry in Rabbit Valley, Colora­do are described. The Mygatt-Moore Quarry has produced thousands of vertebrate fossils from the Brushy Basin Member, with the diplodocid Apatosaurus cf. louisae and the tetanuran Allosaurus fragilis dominat­ing the assemblage. Additionally, remains of another diplodocid, Diplodocus sp., have been found near the quarry within Rabbit Valley. Both specimens in this study preserve eight teeth per alveolar position, as observed through broken surfaces at the gross anatomical level and also through computed tomography (CT) scans. This is inconsistent with the genus Diplodocus sp., which has been previously shown to have a maximum of six teeth per alveolus. The presence of eight replacement teeth per alveolus has previously only been reported in the Cretaceous rebbachisaurid Nigersaurus taqueti, which has been interpreted to have occupied a similar ground-height browsing feeding strategy to both Diplodocus and Apatosaurus. This is the first report of this type of high-count replacement teeth in a diplodocid sauropod from the Morrison Formation. The high number of replacement teeth in a close relative to the contemporaneous Diplodocus provides evidence for niche partitioning among the contemporary ground-height browsing diplodocid sauropods of the Late Jurassic Period in North America.

A preliminary report of the fossil mammals from a new microvertebrate locality in the Upper Jurassic Morrison Formation, Grand County, Utah

The first Mesozoic mammals in North America were discovered in the Morrison Formation during the closing decades of the 19th century, as by-products of dinosaurs quarried by teams led by O.C. Marsh. These tiny fossils served as foundational specimens for our understanding of Mesozoic mammal evolution. There are now nearly 25 mammal-bearing localities known from the Morrison Formation, distributed across the Western Interior from the Black Hills to southern Colorado and west into Utah; the most historically important of these are in Wyoming (e.g., Como Quarry 9). Most Morrison mammals are known by jaws or jaw fragments, and several important Mesozoic groups (e.g., docodonts, dryolestoids, and to a large extent triconodonts and symmetrodonts) were established based on Morrison material, shaping the perception of mammalian diversity on a global scale. Despite heavy sampling of coeval sites elsewhere, the Morrison remains the most systematically diverse (at high taxonomic levels) assemblage of Jurassic mammals in the world. Here, we describe two mammalian specimens and highlight other remains yet to be fully identified from a new microvertebrate locality in the Morrison Formation of eastern Grand County, Utah. The site is positioned low in the Brushy Basin Member and is similar in lithology and stratigraphic level to the famous small vertebrate localities of the Fruita Paleontological Area, located less than 50 km to the northeast. In addition to small archosaurs and squamates, limited excavation to date has yielded at least 20 mammalian specimens representing a minimum of six taxa, several of which are new and quite different from typical Morrison taxa. Preservation is generally excellent and includes partially articulated cranial and postcranial elements of small vertebrates. This new site has great potential to contribute new taxa and more complete morphological data than typical Morrison localities, underscoring the importance of continued field work in the Morrison.

The Upper Jurassic Morrison Formation in north-central New Mexico–Linking Colorado Plateau stratigraphy to the stratigraphy of the High Plains

Most study of the Upper Jurassic Morrison Formation has focused on its spectacular and extensive outcrops on the southern Colorado Plateau. Nevertheless, outcrops of the Morrison Formation extend far off the Colorado Plateau, onto the southern High Plains as far east as western Oklahoma. Outcrops of the Morrison Formation east of and along the eastern flank of the Rio Grande rift in north-central New Mexico (Sandoval, Bernalillo, and San­ta Fe Counties) are geographically intermediate between the Morrison Formation outcrops on the southeastern Colorado Plateau in northwestern New Mexico and on the southern High Plains of eastern New Mexico. Previous lithostratigraphic correlations between the Colorado Plateau and High Plains Morrison Formation outcrops using the north-central New Mexico sections encompassed a geographic gap in outcrop data of about 100 km. New data on previously unstudied Morrison Formation outcrops at Placitas in Sandoval County and south of Lamy in Santa Fe County reduce that gap and significantly add to stratigraphic coverage. At Placitas, the Morrison Formation is about 141 m thick, in the Lamy area it is about 232 m thick, and, at both locations, it consists of the (ascending) sandstone-dominated Salt Wash Member, mudstone-dominated Brushy Basin Member, and sandstone-dominat­ed Jackpile Member. Correlation of Morrison strata across northern New Mexico documents the continuity of the Morrison depositional systems from the Colorado Plateau eastward onto the southern High Plains. Along this transect, there is significant stratigraphic relief on the base of the Salt Wash Member (J-5 unconformity), the base of the Jackpile Member, and the base of the Cretaceous strata that overlie the Morrison Formation (K unconfor­mity). Salt Wash Member deposition was generally by easterly-flowing rivers, and this river system continued well east of the Colorado Plateau. The continuity of the Brushy Basin Member, and its characteristic zeolite-rich clay facies, onto the High Plains suggests that localized depositional models (e.g., “Lake T’oo’dichi’) need to be re-eval­uated. Instead, envisioning Brushy Basin Member deposition on a vast muddy floodplain, with some localized lacustrine and palustrine depocenters, better interprets its distribution and facies.

Evidence for niche partitioning among ground-height browsing sauropods from the Upper Jurassic Morrison Formation of North America

Two tooth-bearing snout fragments from a diplodocid sauropod from the Brushy Basin Member of the Morrison Formation (Upper Jurassic) excavated from the Mygatt-Moore Quarry in Rabbit Valley, Colora­do are described. The Mygatt-Moore Quarry has produced thousands of vertebrate fossils from the Brushy Basin Member, with the diplodocid Apatosaurus cf. louisae and the tetanuran Allosaurus fragilis dominat­ing the assemblage. Additionally, remains of another diplodocid, Diplodocus sp., have been found near the quarry within Rabbit Valley. Both specimens in this study preserve eight teeth per alveolar position, as observed through broken surfaces at the gross anatomical level and also through computed tomography (CT) scans. This is inconsistent with the genus Diplodocus sp., which has been previously shown to have a maximum of six teeth per alveolus. The presence of eight replacement teeth per alveolus has previously only been reported in the Cretaceous rebbachisaurid Nigersaurus taqueti, which has been interpreted to have occupied a similar ground-height browsing feeding strategy to both Diplodocus and Apatosaurus. This is the first report of this type of high-count replacement teeth in a diplodocid sauropod from the Morrison Formation. The high number of replacement teeth in a close relative to the contemporaneous Diplodocus provides evidence for niche partitioning among the contemporary ground-height browsing diplodocid sauropods of the Late Jurassic Period in North America.

Paleontology, taphonomy, and sedimentology of the Mygatt-Moore Quarry, a large dinosaur bonebed in the Morrison Formation, western Colorado—Implications for Upper Jurassic dinosaur preservation modes

The Mygatt-Moore Quarry is a deposit of several thousand dinosaur bones in the Brushy Basin Member of the Morrison Formation in western Colorado. The site has been worked for more than 30 years and nearly 2400 mapped specimens have been collected. This study gathered data about the quarry from many sources to investigate the origin of the deposit. The Mygatt-Moore Quarry appears to be an attritional deposit of a relatively restricted diversity of dinosaurs, with few other non-dinosaurian taxa, that accumulated in a vernal pool deposit in an overbank setting. Bone modification was mostly by corrosion and breakage by trampling; scavenging was abundant. The paleofauna is dominated by Allosaurus and Apatosaurus (MNI and NIS), with the polacanthid ankylosaur Mymoorapelta less common. The matrix of the main quarry layer includes abundant carbonized fragments of plant material, and the mud during the time of deposition may have been often at least damp and occasionally acidic and dysoxic. The Cleveland-Lloyd Dinosaur Quarry is a close correlate of the Mygatt-Moore Quarry in terms of lithology and taphonomy, but demonstrates significant differences upon close inspection of matrix details and bone modification. Large quarries of fine-grained facies in the Morrison Formation possess a very different preservation mode as well as different taxon and relative abundance profiles from those in coarser sediments, which suggests that more may be learned in the future from taphofacies study of large quarries in mudstone beds.

ORIGIN OF THE CARNEGIE QUARRY SANDSTONE (MORRISON FORMATION, JURASSIC) AT DINOSAUR NATIONAL MONUMENT, JENSEN, UTAH

The Carnegie Quarry at Dinosaur National Monument was excavated in the Quarry Sandstone bed of the Brushy Basin Member of the Morrison Formation (Jurassic). The Quarry Sandstone is a discontinuous, lenticular, heterolithic unit that rests upon drab, greenish-gray claystones and is capped by reddish-brown mudstones. The underlying drab claystones are interpreted as being deposited in an overbank interfluvial wetland or lacustrine environment. By tracing the sandstone and studying detailed measured stratigraphic sections along its 1.8-km outcrop belt, this interval can be shown to be a complex of intergrading sandstone bodies and mudstone units. The Quarry Sandstone interval is reinterpreted as being deposited within a crevasse splay complex. The lowest part of this deposit is a laterally continuous, tabular unit composed of polymictic sandstone and granule conglomerate that directly overlies lacustrine claystones. This unit contains re-sedimented carbonate clasts from the surrounding Brushy Basin, displays a gradational to sharp basal contact, and is interpreted as the initial crevasse splay deposit. Overlying the tabular basal unit are numerous, laterally continuous, upward-coarsening sandstone bodies interpreted as distributary mouth bars. The interpreted distributary mouth bar deposits are cut by lenticular, trough cross-bedded, upward fining sandstone units exhibiting erosional bases. These sandstone lenses are interpreted as distributary channels within the crevasse splay. They, in turn, are overlain by discontinuous sandstone bodies that interfinger with, and are overlain by reddish brown and variegated, rooted mudstones containing calcic paleosols. The Carnegie dinosaur quarry was excavated in one of the main distributary channels of this splay complex.

A preliminary report of the fossil mammals from a new microvertebrate locality in the Upper Jurassic Morrison Formation, Grand County, Utah

The first Mesozoic mammals in North America were discovered in the Morrison Formation during the closing decades of the 19th century, as by-products of dinosaurs quarried by teams led by O.C. Marsh. These tiny fossils served as foundational specimens for our understanding of Mesozoic mammal evolution. There are now nearly 25 mammal-bearing localities known from the Morrison Formation, distributed across the Western Interior from the Black Hills to southern Colorado and west into Utah; the most historically important of these are in Wyoming (e.g., Como Quarry 9). Most Morrison mammals are known by jaws or jaw fragments, and several important Mesozoic groups (e.g., docodonts, dryolestoids, and to a large extent triconodonts and symmetrodonts) were established based on Morrison material, shaping the perception of mammalian diversity on a global scale. Despite heavy sampling of coeval sites elsewhere, the Morrison remains the most systematically diverse (at high taxonomic levels) assemblage of Jurassic mammals in the world. Here, we describe two mammalian specimens and highlight other remains yet to be fully identified from a new microvertebrate locality in the Morrison Formation of eastern Grand County, Utah. The site is positioned low in the Brushy Basin Member and is similar in lithology and stratigraphic level to the famous small vertebrate localities of the Fruita Paleontological Area, located less than 50 km to the northeast. In addition to small archosaurs and squamates, limited excavation to date has yielded at least 20 mammalian specimens representing a minimum of six taxa, several of which are new and quite different from typical Morrison taxa. Preservation is generally excellent and includes partially articulated cranial and postcranial elements of small vertebrates. This new site has great potential to contribute new taxa and more complete morphological data than typical Morrison localities, underscoring the importance of continued field work in the Morrison.