Anthropogenic Pollen Research Articles

BRAIN - Holocene archaeo-data for assessing plant-cultural diversity in Italy and other Mediterranean regions

In the field of botany applied to archaeological and palaeoecological studies, the multi- and inter-disciplinary nature of this research produces a lack of data sharing and scattered articles in the specialty literature or in national and international journals. The vast production of archaeobotany and palynology data makes it necessary to develop a tool for the availability, accessibility, and dissemination of existing research. Many databases exist on palaeoecology, archaeobotany or pollen data. There are no collections focused on archaeological sites and human-induced environments and centred on Southern Europe and the Mediterranean. BRAIN - Botanical Records of Archaeobotany Italian Network is the first database listing sites from which all types of plant records are available in Italy and nearby Mediterranean regions. BRAIN represents the largest integrated collection of archaeo/palaeo-botanical data and a range of descriptive information that makes data recovery FAIR ready. This unique network hosts data on the availability of anthropogenic pollen, palynomorphs and plant macroremains in the same database, and experts of different research fields may contribute to it.

The Establishment of the Agricultural Landscape of Central Sicily Between the Middle Neolithic and the Beginning of the Iron Age

The possible co-variation of human occupation and vegetation from the Middle Neolithic to the beginning of the Iron Age (7.5–2.8 ka BP) in Central Sicily in the context of the central Mediterranean between Middle and Late Holocene are analysed in this paper to provide new insights on Sicilian prehistoric demography. The demographic and economic trends during these millennia were reconstructed using archaeological, Accelerator Mass Spectrometry 14C dates, palynological, archaeobotanical, and zooarchaeological data from the northern, central, and southern sectors of Central Sicily through a diachronic comparison with variation in Arboreal Pollen, Anthropogenic Pollen Indicators, Olea-Juglans-Castanea pollen, microcharcoals, and Sporormiella from four pollen cores from sites in different ecosystems. A very significant spread of farming activities was found at the end of the Neolithic, together with an apparent demographic gap during the Middle Copper Age, and the emergence of agricultural landscapes at the end of the Copper Age associated with a striking increase in population. A combination of cultural and climatic changes during the late phase of the Bronze Age resulted in a subsequent overall decrease in population.

Reconstruction of Holocene precipitation patterns and vegetation evolution in the North China Plain: Deciphering the relative influence of climate and anthropogenic forcing

In this paper, we investigate Holocene precipitation changes in northern China to improve the understanding of the East Asian summer monsoon (EASM) and provide an environmental context for the development of Chinese civilisation. Our study is based on an analysis of 260 pollen samples from Holocene lacustrine sediments at Ningjinpo in northern China, and employs the biomisation method to reconstruct regional vegetation types and vegetation changes, and to decipher their driving factors, by incorporating regional climatic data and information on human activities. Results indicate that, in the early Holocene (10,000–8100 cal yr B.P.), regional climate was dry, with annual precipitation of 470 mm. In the middle Holocene (8100–3000 cal yr B.P.), a rapid increase was observed in precipitation, reaching 580 mm. During the late Holocene (2600–600 cal yr B.P.), climate became drier again, with annual precipitation decreasing to 480 mm. Between 10,000–600 cal yr B.P., vegetation exhibited distinct successional patterns dominantly influenced by climate, transitioning from meadow-steppe to forest, and then back to meadow-steppe. After 600 cal yr B.P., vegetation succession decoupled from climate change, with human activities emerging as the primary driving force, as indicated by the substantial increase in anthropogenic pollen and charcoal concentrations. In the future, under the combined influence of global warming and intensified human activity, the implementation of scientific environmental planning measures will become increasingly important for the restoration and sustainable development of regional vegetation ecosystems.

Anthropogenic pollen indicators: Global food plants and Latin American human indicators in the pollen record

Pollen-based evidence of human presence is crucial for reconstructing human history. However, information on the morphology of pollen grains of global food plants and regional pollen-based human indicators is scattered in the literature, leading to the risk of overlooking important evidence of human presence. To address this issue, we first compiled a comprehensive overview of 354 major food plants worldwide, creating a paleoecology-friendly format that includes their family, vernacular name, earliest known use, environmental preference, and geographical region. Moreover, we identified the sources of illustrations of their pollen grains for 209 out of 273 different genera of globally relevant food plants in 10 selected pollen atlases. Secondly, we compiled all human indicators from pollen-based paleoecological literature in Latin America (based on 750 references), providing an overview of 212 single-pollen type indicators and identified 95 crucial combinations of pollen types as “human indices”, and their corresponding references. Our review datasets aids in distilling human evidence from numerous fossil pollen records worldwide.

Paleoecological data complete historical and archaeological archives: Human–environment nexus at the fluvio maritime city port of Chilia (Licostomo; Northern Danube delta) since the 14th century

AbstractThis study reconstructs the paleoecological settings of Chilia/Licostomo, the most important Genoese colony in the Danube delta. Our paleoenvironmental data (i.e., sedimentology, pollen, charcoal, radiocarbon ages) corroborate historical and archaeological information over the last seven centuries to document the habitation at Chilia and its military and socioeconomic activity during ca. 14th–19th centuries A.D. Palynological data show the continuous presence of humans, with anthropogenic pollen index calculation (API) values between 2% and 16% during the last five centuries. The variability of API, corroborated with the major peaks of the charcoal concentration, matches the documented military conflicts and population changes in the area: the Ottoman conquest of Dobrugea (15th century A.D.), the Zaporozhians Cossaks raids (17th century A.D.), and the Russian–Ottoman wars (18th–19th centuries A.D.). The presence of coprophilous fungi fits the Ottoman tradition of animal husbandry, whereas their appearance after the Zaporozhians Cossacks raids and their good correlation with open land vegetation suggest that the Chilia loess island was temporally used as a refuge from multiple menaces (e.g., wars, administrative crisis). These paleoecological data, together with the rare archaeological findings, exclude human habitation of the Chilia Plain earlier than the documented arrival of the Lipovans during the late 18th century A.D. The critical assessment of the correspondences between the paleoecological and historical data shows the relevance of the approach in deltaic environments, when applied to lagoon/lacustrine sediments, yet often limited to chronologically disparate, decennial‐ to multicentennial‐scale temporal frameworks.

Human-Impact Gradients through Anthropogenic Pollen Indicators in a Mediterranean Mosaic Landscape (Balearic Islands)

This paper proposes new anthropogenic pollen indicators for the Balearic Islands and attempts to assess gradients of human impact on vegetation in Mediterranean islands. A combination of modern pollen analogue studies, complemented by phytosociological descriptions and ordination techniques using quantitative and presence/absence data was used. Redundancy analysis allowed us to evaluate the relationships between pollen types and significant environmental variables and propose regional (e.g., Centaurea, Rubus, Plantago lanceolata-t) and local/microregional anthropogenic pollen indicators (e.g., Cerealia, Poygonum aviculare, Matricaria-t). Additionally, an anthropogenic index score (AIS) for each sampled location was calculated to correlate each pollen type to a specific degree of human impact: (a) low (e.g., Cerastium-t, Erica arborea-t, Cistus albidus), (b) moderate (e.g, Sinapis-t, Sanguisorba minor-t, Plantago bellardii-t), (c) high (e.g., Papaveraceae undiff., Dipsacaceae, Secale-t). This paper contributes to a further understanding of land-use dynamics and to defining the degree of impact, which is especially necessary to assess colonization and anthropization rhythms in Mediterranean island environments.

Flora-vegetation history and land use in Medieval Tuscany: The palynological evidence of a local biodiversity heritage

This paper introduces the high resolution palynological analyses carried out on samples from four cores drilled in the coastal plains of the Cornia and Pecora rivers, southern Tuscany, Italy. This study provides new information on the landscape transformations that the area has undergone over the past 7500 years, with focus on Medieval times when settlement patterns and land use contributed the onset of the current landscape. The study area, considered one of the most important early centres of the European civilization, has been fully investigated by the ERC funded nEU-Med project through an interdisciplinary approach combining archaeological, geomorphological, chemical, and archaeobotanical analyses. Environmental features and resource availability were explored through pollen markers of natural environments and human activities suggesting similarities and dissimilarities between the two valleys. In the Cornia Valley, the high diversity and percentages of woody taxa is connected to agrarian exploitation with arboriculture, while the Pecora Valley had a mostly open landscape, with lower diversity and higher percentages of anthropogenic pollen indicators. In both valleys, the land was mainly exploited for grazing resources: the high presence of pasture indicators suggests that domesticated animals should have been an important local resource especially in Medieval times. Pastoral/breeding activities fit into a larger context of management of the territory with specific vocation (namely, salt and iron in these two valleys). The detailed palynological analysis and the comparison between the two valleys highlight the local character of these activities, probably carried out in a collateral way by the peasant communities between the 7th-12th centuries AD.

Sedimentological and palynological records since 10 ka BP along a proximal-distal gradient on the Armorican shelf (NW France)

New findings acquired in Armorican shelf (core MD08-3204 CQ: Bay of Quiberon and core VK03-58bis: South Glénan islands) depict Holocene paleoenvironmental changes since 10 ka BP through a multi-proxy dataset including sedimentological and palynological data. First, grain-size analyses and AMS-14C dates show a common sedimentary history for both study cores. The slowdown of the relative sea level (RSL) rise was accompanied by a drop of the sedimentation rates between ∼8.3 and 5.7 ka BP; the rates had been relatively higher at the onset of the Holocene. This interval led to the establishment of a shell-condensed level, identified in core VK03-58bis by the “Turritella layer” and interpreted as a marker for the Maximum Flooding Surface. Palynological data (pollen and dinoflagellate cyst assemblages) acquired in core MD08-3204 CQ argue for an amplification of the fluvial influence since 5.7 ka BP; the establishment of the Highstand System Tract (i.e., mixed marine and fluviatile influences on the platform) then accompanied the slowdown of the RSL rise rates. On the shelf, the amplification of Anthropogenic Pollen Indicators (API) has been better detected since 4.2 ka BP, due not only to the increase of human impact but also due to a stronger fluvial influence on the shelf during the Late Holocene. Palynological data, recorded on the 8.5–8.3 ka BP interval along an inshore-offshore gradient, also demonstrate the complexity of the palynological signals such as i) the fluvial influence that promotes some pollen taxa (Alnus and Corylus) from proximal areas and ii) the macro-regionalization of palynomorph sources in distal cores. In addition, the comparison of palynological tracers, including API, over the last 7 kyrs BP, with South Brittany coastal and mid-shelf sites subject to northern vs. southern Loire catchment areas, shows a major hydro-climatic effect on the reconstructed palynological signals. Strengthened subpolar gyre (SPG) dynamics, combined with recurrent positive North Atlantic Oscillation (NAO) configurations, appear responsible for increased winter precipitation and fluvial discharge over northern Europe, as in Brittany. Conversely, weakened SPG intervals, associated with recurrent negative NAO-like modes, are characterized by intensified winter fluvial discharge over southern Europe. Interestingly, at an infra-orbital timescale, we record major peaks of API during periods of strengthened (/weakened) SPG dynamics in sites subject to detritic-organic matter inputs from Brittany (/Loire) watersheds.

Agricultural practices in India during the Holocene: A pollen view point and a critical appraisal

Pollen assemblages provide valuable insights into the beginning of cereal-based agricultural practices and the transition from a hunting and gathering to a sedentary and food-producing way of life. Anthropogenic pollen indicators (APIs) and their precise identification, with respect to taxonomic resolution, can help to document the history of agricultural development, pastoral activities and human-induced land-use changes, particularly for the Holocene Epoch. Moreover, careful selection of pollen types, and/or indices, established for a particular region, are useful for obtaining meaningful reconstructions of anthropogenic activities through time. Specific pollen-markers have been used to deduce the inception of agriculture and the impact of anthropogenic activities on the landscape, from the different regions of the world. For India, Cerealia, and other cultural plant pollen taxa, such as Amaranthaceae, Caryophyllaceae, Brassicaceae, Polygonaceae, Artemisia, Cannabis sativa, Alternanthera, Urtica, Rumex and Borreria have been used as marker pollen types for indicating agricultural practices and other anthropogenic activities. In this communication, we have made an attempt to trace the advent and intensification of agricultural practices in the varied physiographic regions of the Indian sub-continent, on the basis of the available palynological records. We have further critically analysed the plausibility of agricultural practices and other human activities during the Late Pleistocene/Pre and Early Holocene with respect to the Indian scenario.

Managing wilderness? Holocene-scale, human-related disturbance dynamics as revealed in a remote, forested area in the Czech Republic

In the lowlands and uplands of Central Europe, which were inhabited continuously from the very start of the Holocene to the present times, it is difficult to find territories suitable for investigation of natural baselines. For this reason, we picked the complicated rocky terrain of one upland area in NE Bohemia called Adrspach because, based on the absence of archeological finds, it was supposed to have never been deforested or managed by people. The remote and inhospitable character of this particular area further encouraged this assumption. To our great surprise, however, high-resolution pollen analyses, supplemented by analyses of non-pollen palynomorphs and microscopic charcoals reveals that the local forest ecosystem had a dynamic development over entire Holocene. We were able to correlate this high-resolution understanding of vegetation successions with repeated fire disturbances. Was this fire disturbance dynamic natural? Subsequent archeological exploration and excavation in the area brought unexpected evidence, pointing to rather continuous human presence throughout most of the Holocene. From the Early Mesolithic to the Late Neolithic, available evidence suggests a hunter-gatherer mode of resource management. From the start of the Late-Holocene (ca 4 ka BP), the occurrence of coprophilous fungal spores and secondary anthropogenic pollen indicators suggest this area was impacted by recurrent domestic animal grazing. Testing this approach also in other remote forested areas of Central Europe, we argue, can have far-reaching implications for understanding long term human-environment agency by transforming our understanding of alternative subsistence and land use strategies during prehistory. At the same time, this can significantly alter existing concepts used in Central European nature conservation strategies, which tend to be based on an underlying assumption that our work challenges – the survival of little impacted wilderness at the Holocene scale.

Late-Holocene paleoenvironmental and land-use changes in Western Greece based on a sediment record from Klisova lagoon

In this study, we present the findings of a sediment core retrieved from Klisova lagoon, Western Greece, an area with a long record of documented human presence. The recovered deposits were subjected to sedimentological, XRF, and micropaleontological analyses. For the last 4700 cal BP, the freshwater influx, the progradation of the Evinos river delta and related geomorphological changes control the environmental conditions in the lagoon. Considering the centennial temporal resolution of our analyses, small offsets of c.a. 50 years due to lack of regional reservoir correction do not considerably impact the reported radiocarbon ages. Prior to 4000 cal BP, a relatively shallow water depth, significant terrestrial/freshwater input and increased weathering in the lagoon area are inferred. Elemental proxies and increased dinoflagellate cyst and foraminiferal abundances, which indicate marine conditions with prominent freshwater influxes, point to the gradual deepening of the lagoon up to 2000 cal BP. The marine and freshwater condition equilibrium sets at 1300 cal BP, with the lagoonal system reaching its present state. Maxima of anthropogenic pollen indicators during the Mycenaean (3200 cal BP), Hellenistic (2200 cal BP), and Late Byzantine (800 cal BP) periods suggest intervals of increased anthropogenic activities in the area.

Influence of taxonomic resolution on the value of anthropogenic pollen indicators

The taxonomic resolution of palynological identification is determined by morphological criteria that are used to define pollen types. Different levels of taxonomic resolution are reached in palynology, depending on several factors such as the analyst’s expertise, the palynological school, the aim of the study, the preservation of the pollen grains, the reference collections and the microscope facilities. Previous research has suggested that attaining pollen records with high taxonomic resolution is important to reconstruct correctly past land use and human impact. This is in turn central to disentangling past human activities from other drivers of long-term vegetation dynamics such as natural disturbance or climate variability. In this study, we assess the impact of taxonomic resolution on the indicative capacity of anthropogenic pollen types. To achieve this, we attribute the pollen types of sixteen sedimentary records, located along a latitudinal gradient spanning from Switzerland to Italy, to three levels of taxonomic resolution previously proposed at the European scale. Our results show that higher taxonomic resolution improves the identification of human impact by enhancing the indicative power of important pollen indicators widely used in the research field. Our results may contribute to the improvement of palynological reconstructions of land use and human impact by identifying key pollen types whose determination requires particular attention.

Climate–human–environment interactions in the middle Yangtze Basin (central China) during the middle Holocene, based on pollen and geochemical records from the Sanfangwan Site

The Shijiahe Culture (ca. 4800–4100 yr BP), which originated in the middle Yangtze Basin, was a milestone of prehistoric Chinese civilization. To reveal the interactions between climatic changes and human activities during this time interval, the geochronology and multiple proxies (pollen and heavy metals) of samples from a 220-cm-deep outer wall at the Sanfangwan Site (a typical site of the Shijiahe Culture) were analyzed. Radiocarbon dating indicates that the age range of the Sanfangwan Site is ca. 4800 to 4100 cal. yr BP. From ca. 4800 to 4600 cal. yr BP (the early Shijiahe Culture), pollen assemblages were dominated by Gramineae and an anthropogenic plant community (e.g. Ranunculaceae, Chenopodiaceae, Labiatae, and Cruciferae), indicating an expansion in rice agriculture during this interval. High metal concentrations in the same interval indicate a large flux of metals from smelting and farming activities. Between ca. 4600 and 4100 cal. yr BP (the middle and late Shijiahe Culture), decreases in Gramineae and anthropogenic pollen mirrored the recession of farming. Concurrent decreases in heavy metals may be attributable to a decline in the smelting industry. Three episodes of high pollen concentrations and visible increases in hygrophyte pollen, Pinus pollen, and fern spores at 4600–4550, 4450–4350, and 4200–4100 cal. yr BP probably indicate the occurrence of several flooding events during the middle and late Shijiahe Culture, resulting in the migration of local inhabitants. This study highlights the driving effect of flood events on the collapse of the Shijiahe Culture.

The prehistory and early history of the Šumava Mountains (Czech Republic) as seen through anthropogenic pollen indicators and charcoal data

In the lower forested mountain ranges of Europe, human impact on nature is usually confined to the written history of the Middle Ages. Our research in the Šumava mountains aims to specify the nature and intensity of human impact on vegetation, especially during agricultural prehistory. We use results from a multidisciplinary study of the unique La Tène archaeological site in the Sklářské Valley (elevation 802 m a.s.l.) and from a pollen and charcoal record 60 m away. With knowledge of this reference site we focus on the meaning of anthropogenic pollen indicators in 13 other pollen sites from central Šumava. From ca 3300 cal yr BP we detect an increase in NAP, Betula, Pinus and secondary anthropogenic indicators in pollen records – vegetation changes of anthropogenic origin. Charcoal records show a significant peak around 3200 cal yr BP. We found weak anthropogenic influence on the La Tène archaeological site in the Sklářské Valley, and much stronger anthropogenic pollen signals at other pollen sites dated to the Late Bronze and Iron Ages. Some of these sites are situated on trade routes which have been known since Medieval times but which most likely have much older origins. During prehistory, pollen data reveal no specific human activity such as pasturing or arable farming but reflect small-scale disturbances that supported growth of Betula and Pinus and an abundance of herbs. Such human impact could be connected primarily to activities along trade routes and to hunting, but other factors cannot be excluded.

A critical assessment of human-impact indices based on anthropogenic pollen indicators

Anthropogenic pollen indicators in pollen records are an established tool for reconstructing the history of human impacts on vegetation and landscapes. They are also used to disentangle the influence of human activities and climatic variability on ecosystems. The comprehensive anthropogenic pollen-indicator approach developed by Behre (1981) has been widely used, including beyond its original geographical scope of Central and Western Europe. Uncritical adoption of this approach for other areas is risky because adventives (plants introduced with agriculture) in Central Europe can be apophytes (native plants favoured by human disturbances) in other regions. This problem can be addressed by identifying region-specific, anthropogenic-indicator pollen types and/or developing region-specific, human-impact indices from pollen assemblages. However, understanding of regional variation in the timing and intensity of human impacts is limited by the lack of standardization, validation and intercomparison of such regional approaches. Here we review the most common European anthropogenic pollen-indicator approaches to assess their performance at six sites spanning a continental gradient over the boreal, temperate and Mediterranean biomes. Specifically, we evaluate the human-indicator approaches by using independent archaeological evidence and models. We present new insights into how these methodologies can assist in the interpretation of pollen records as well as into how a careful selection of pollen types and/or indices according to the specific geographical scope of each study is key to obtain meaningful reconstructions of anthropogenic activity through time. The evaluated approaches generally perform better in the regions for which they were developed. However, we find marked differences in their capacity to identify human impact, while some approaches do not perform well even in the regions for which they were developed, others might be used, with due caution, outside their original areas or biomes. We conclude that alongside the increasing wealth of pollen datasets a need to develop novel tools may assist numeric human impact reconstructions.

Climate and vegetation dynamics of the Northern Apennines (Italy) during the Late Pleistocene and Holocene

This study reconstructs the regional vegetation and climate dynamics between the upper Late Pleistocene and Holocene around Pian del Lago, a coastal mountain marshland located at 831 m asl in western Liguria (NW-Italy), based on the pollen analysis of a 13 m-long sediment core. The record provided a unique opportunity to study a poorly documented period in northern Italy and across many parts of southwestern Europe. We propose an event stratigraphy based upon the identification of seven interstadials (NAI-7 to NAI-1) spanning the upper Late Pleistocene. The correlation with other terrestrial records in Italy, and with Mediterranean marine sequences and the Greenland ice cores, permitted a coherent reconstruction of main environmental changes from >∼43,000 cal. BP. Significantly, the pollen record indicates the persistence of a mesophilous mountain vegetation cover, mainly composed of Quercus (deciduous and evergreen), Abies, Fagus and Alnus over the whole time period recorded. At the Last Glacial Maximum (LGM) and during the Late Würm Lateglacial, despite the presence of steppic vegetation composed of Artemisia, woodlands dominated by Pinus, with Abies, Picea, Fagus, Alnus and Betula are present. This forest composition provides an important insight into the history of Picea in southern Europe and Late Pleistocene refugia for mesophilous species. During the Early Holocene, Pinus is first replaced by Abies and then by deciduous Quercus and mixed temperate species as the dominant forest component. Both arboreal and herbaceous anthropogenic pollen indicators only make their appearance during the Late Holocene, attesting to the increasing importance of human activities.

Mediterranean landscape change during the Holocene: Synthesis, comparison and regional trends in population, land cover and climate

This synthesis paper offers a comparative perspective on how seven different Mediterranean regions, from Iberia and Morocco to the Levant, have been transformed by human and natural agencies during the past 10 millennia. It draws on a range of data sources: notably (1) archaeological site surveys ( n = 32,000) and 14C dates ( n = 12,000) as proxies for long-term population change, (2) pollen records as a proxy for past vegetation and land cover ( n = 253) and (3) proxies, such as stable isotopes, from lake, cave and marine records as indicators of hydro-climate ( n = 47). Where possible, these data sets have been made spatially and temporally congruent in order to examine relationships between them statistically and graphically. Data have been aggregated or averaged for each region/sub-region and put into 200-year time windows. Archaeo-demographic data show a clear increase at the start of Neolithic farming, followed by a series of regionally asynchronous fluctuations in population, prior to a pan-Mediterranean Roman settlement maximum. Pollen data indicate a late-Holocene decline in %Arboreal Pollen in those regions that were initially well wooded, but not in drier regions of the southern/eastern Mediterranean. Overall, the clearest palynological proxy for human land cover change is provided by the OJCV (tree crop) index. The cultivation of these trees in the eastern Mediterranean after 6500 cal. yr BP may have been an adaptive response to mid-Holocene climatic desiccation. These anthropogenic pollen indicators correlate more closely with trends in population than with regional hydro-climatic z scores, implying that they reflect primarily human activities. During the mid Holocene, most Mediterranean landscapes were transformed by a combination of climate and rural land use, but after ~3500 cal. yr BP, human actions became increasingly dominant in determining land cover. During the past 1500 years, the dominant landscape trajectory in the eastern Mediterranean was markedly different to that in the central/western Mediterranean.

Pollen-inferred regional vegetation patterns and demographic change in Southern Anatolia through the Holocene

Southern Anatolia is a highly significant area within the Mediterranean, particularly in terms of understanding how agriculture moved into Europe from neighbouring regions. This study uses pollen, palaeoclimate and archaeological evidence to investigate the relationships between demography and vegetation change, and to explore how the development of agriculture varied spatially. Data from 21 fossil pollen records have been transformed into forested, parkland and open vegetation types using cluster analysis. Patterns of change have been explored using non-metric multidimensional scaling (nMDS) and through analysis of indicator groups, such as an Anthropogenic Pollen Index, and Simpson’s Diversity. Settlement data, which indicate population densities, and summed radiocarbon dates for archaeological sites have been used as a proxy for demographic change. The pollen and archaeological records confirm that farming can be detected earlier in Anatolia in comparison with many other parts of the Mediterranean. Dynamics of change in grazing indicators and the OJCV ( Olea, Juglans, Castanea and Vitis) index for cultivated trees appear to match cycles of population expansion and decline. Vegetation and land use change is also influenced by other factors, such as climate change. Investigating the early impacts of anthropogenic activities (e.g. woodcutting, animal herding, the use of fire and agriculture) is key to understanding how societies have modified the environment since the mid–late Holocene, despite the capacity of ecological systems to absorb recurrent disturbances. The results of this study suggest that shifting human population dynamics played an important role in shaping land cover in central and southern Anatolia.

Human demography changes in Morocco and environmental imprint during the Holocene

The aim of this work is to reconstruct the periods of growth and decline of human populations in Morocco and their potential impacts on the landscape over the past 10,000 years. In order to estimate the trends in the human population size between 10,000 and 3000 years ago, we used a summed probability distribution (SPD) of radiocarbon dates from a wide range of archaeological sites throughout Morocco. Landscape changes were identified and quantified from a dataset of fossil pollen records. Different anthropogenic pollen markers, as well as natural vegetation groups and taxonomic richness were used to analyse the relationship between long-term trends in human population expansion or regression and type of impact on the landscape. The sub-regions of Morocco have different topographies and climates, which have either favoured or prevented the establishment and/or spread of human populations. In order to identify the areas most significantly impacted by humans and the timing of such impacts, we have reconstructed and compared the same past anthropogenic and landscape proxies along with the population trends within the lowlands and mountainous areas. The lowlands were more strongly impacted earlier in the Holocene than the mountainous areas. Anthropogenic markers indicate that farming expanded in the lowlands during the first major expansion of human populations between ca. 7200 and 6700 cal. yr BP at the start of the Neolithic period. In the Atlas and Rif Mountains, anthropogenic impact is not clearly detectable in any of these areas before 4000 cal. BP.

The 4.2 ka event in the vegetation record of the central Mediterranean

Abstract. In this paper, the variation in forest cover in the central Mediterranean region, reflected by percentage changes in the arboreal pollen record, has been examined in relation to the 4.2 ka event. A total of 36 well-dated and detailed pollen records from latitudes between 45 and 36∘ N were selected and their vegetation dynamics between 5 and 3 ka examined in relation to the physiographic and climatic features of the study area and to the influence of human activity on past vegetation, as suggested by anthropogenic pollen indicators. We have found that the sites located between 43 and 45∘ N do not show any significant vegetation change in correspondence with the 4.2 ka event. Several sites located on the Italian Peninsula between 39 and 43∘ N show a marked opening of the forest, suggesting a vegetation response to the climate instability of the 4.2 ka event. Between 36 and 39∘ N, a forest decline is always visible around 4.2 ka, and in some cases it is dramatic. This indicates that this region was severely affected by a climate change towards arid conditions that lasted a few hundred years and was followed by a recovery of forest vegetation in the Middle Bronze Age. Human activity, especially intense in southern Italy, may have been favored by this natural opening of vegetation. In Sardinia and Corsica, no clear change in vegetation is observed at the same time. We suggest that during the 4.2 ka event southern Italy and Tunisia were under the prevalent influence of a north African climate system characterized by a persistent high-pressure cell.