Analysis Of Macroremains Research Articles

The first comprehensive macroremains analysis of edible plants from Vichama, Peru (1800–1500 bce)

The first comprehensive macroremains analysis of edible plants from Vichama, Peru (1800–1500 bce)

The medieval croft plužina field system in a mountain region of central Europe: The interdisciplinary record of the earthen field boundaries in Debrné, Czechia

AbstractThe integration of archaeological, historical and geoarchaeological records represents a significant contribution to research into the medieval landscape. This study focuses on the medieval field system in the deserted village of Debrné, located in northeastern Bohemia, Czechia. The village features a well‐preserved croft plužina field system, a typical historical landscape of central Europe. The main and most visible elements of the field system are the earthen field boundaries, which were the focus of the geoarchaeological investigations. Archaeological excavations in trench S1 revealed a collection of larger stones at a depth of 1 m, potentially serving a drainage function akin to the observed plužina. Additionally, a boulder paving, identified as a remnant of a path between fields, provided insights into the historical use of the area. In trench S2, positioned closer to the village's core, layers with increasing stone content were recorded at a depth of 130 cm. However, the drainage structure observed in trench S1 was not replicated. The dating of earthen field boundaries indicated the creation of the terrace in the second half of the 16th century in trench S1. In trench S2, radiocarbon dating at a depth of 70 to 80 cm ranged from 1025 to 1175 A.D., predating the first written source about Debrné. Optically stimulated luminescence (OSL) dating in trench S2 suggested exposure to sunlight during the third century A.D. Archaeobotanical analysis of charred macroremains from trench S1 revealed 236 plant macroremains belonging to approximately 20 taxa, showcasing wild‐growing, collected useful species such as raspberry, blackberry and elderberry. Trench S2 yielded 23 plant macroremains belonging to 11 taxa, with similar species as trench S1. Pedological and micromorphological examinations displayed distinct layering in both trenches, showing up to six layers. Micromorphological analysis unveiled vuggy microstructures, varying grain sizes and elemental patterns, shedding light on different periods of occupation. Multidisciplinary investigations of the buried soils forming the earthen field boundaries discovered that the original soil cover comprised automorphic lixisols and cambisols, which also form under present conditions. These results underscore the importance of integrating pedological, geoarchaeological, archaeobotanical and physical data to comprehend the intricate nature of anthropogenic landscape changes.

How animal dung can help to reconstruct past forest use: a late Neolithic case study from the Mooswinkel pile dwelling (Austria)

Animal dung analyses are a useful tool for vegetational studies. Preserved ruminant dung from archaeological layers offers a unique possibility for the reconstruction of past fodder management strategies, and further for studying the impact of fodder acquisition and pasturing on forests. In this case study we investigate the impact of Late Neolithic livestock keeping on the forest around the “Mooswinkel” pile dwelling at the Austrian lake Mondsee through the analysis of botanical macroremains, insect remains as well as microhistological analyses of botanical remains in animal dung. Seasonal plant parts in the dung point out that cattle, goats, and/or sheep were evidently kept inside the settlement during the winter for protection. During the daytime, they were allowed to forage around the settlement. Winter fodder consisted of dried leaf hay, hay from grasses and herbs, male flowers of early blooming bushes, and fresh twigs of evergreen species, such as fir (Abies alba), ivy (Hedera helix), and mistletoe (Viscum album).

Holocene dynamics of a relict moss complex in the Korotovskoe mire (State Complex «Zavidovo», Russia)

The Korotovskoe mire in the National Park «State complex “Zavidovo”» is the only mire massif in the Moscow Region (Russia) where the postglacial relict complex of vascular plants (e.g. Betula humilis, Pedicularis sceptrum-carolinum, Saxifraga hirculus, Trichophorum alpinum) and mosses (e.g. Cinclidium stygium, Drepanocladus trifarius, Hamatocaulis vernicosus, Meesia triquetra, Paludella squarrosa, Scorpidium scorpioides, S. cossonii, Tomentypnum nitens) is maximally saved. This complex is typical for spring fens. We studied the main stages of the mire development during the last 10 000 years using the analysis of macroremains and radiocarbon analysis. It was shown that despite the instability of the water regime on the mire, minerotrophic mosses were found in the most peat layers. This fact confirms the stability of mineral water inflow during the early and middle Holocene. Meesia triquetra and Drepanocladus trifarius live on the mire since the Boreal period. During the Atlantic and Subboreal period (when the mire surface dried out) there was a very slow peat deposition and the mire was covered by trees, together with a decrease in minerotrophic mosses in the vegetation cover. Scorpidium cossonii, Hamatocaulis vernicosus and Drepanocladus sendtnerii emerge during peat deposits in the Subboreal period after the water regime restoration and the returning of swamp conditions. In the last 2000 years, the surface of the mire was covered by Sphagnum mosses. During the meso-oligotrophic stage of the mire development, mosses demanding a rich mineralisation remained in other parts of the mire, where conditions of rich mineral nutrition lasted longer. Limiting factors of this moss complex during the early and middle Holocene were light and level of mire waters, whereas, in the late Holocene, it was oligotrophisation of the upper peat layers because of a rapid deposition of organic matter. The complex of relict minerotrophic mosses currently existing on mires in the centre of European Russia can be considered as relicts of the glacial time, and, at the same time, as relicts of spring fens, which were more widespread in the early and middle Holocene and gradually having lost their importance as a result of autogenic succession. The obtained data indicate the great nature conservation value of the Korotovskoe mire massif.

Holocene regression of the critically endangered species Cladium mariscus (L.) Pohl on Žitný ostrov Island (site Mad, Danubian plain Lowland)

This study presents the palaeo-botanical record from the central part of Žitný Ostrov Island on the Danubian Lowland in Slovakia. The record spans almost the entire Holocene period and provides evidence that the wetland species Cladium mariscus (Cyperaceae) was present in this region as late as the Greenlandian period. In addition, the combination of C. mariscus and Carex elata beds were most likely an important part of the littoral zone vegetation during that period. Our investigations are based on plant macro-remain analysis, radiocarbon dating and physical sedimentary analysis of the former Danube side channel. The palaeo-channel infill active in the Late Pleistocene reflects complete hydroseral succession of a terrestrialised palaeo-lake. The earlier of two possible hiatuses in the fossil record was likely associated with a disturbance which caused Cladium mariscus to disappear from the local plant spectra. The most likely drivers of C. mariscus’ decline are a climate change of 8.6 cal ka BP associated with water fluctuations and the local habitat alteration to an eutrophic environment. These are discussed herein together with the isolated scattered populations surviving in the surrounding micro-region throughout the Meghalayan period. Finally, there are only two known remaining C. mariscus sites in the Žitný Ostrov Island area, and the key modern driving factor was the dramatic water regime changes in 19–20th Century caused by anthropogenic impact. These were caused by the construction of drainage canal systems and the transition from alluvial meadows to agricultural land.

Eemian (MIS 5e) climate oscillations based on palaeobotanical analysis from the Beckentin profile (NE Germany)

Climate instability in the Eemian Interglacial is gaining more and more evidence. One of such oscillations in the Beckentin record is indicated by the disappearance of the Cladocera and changes in macroremains at the beginning of E4a (Corylus phase). They confirm the shallowing of the reservoir, probably caused by climate changes, but not reflected in the palynological record. However, a significant oscillation observed in the first half of the E5 (Carpinus phase), ~5860 years after the boreal forests subsided and ~6290 years after the onset of interglacial forestation, suggests that, although not being an analogue of the OHO (Older Holsteinian Oscillations) in MIS 11c and the 8.2 ka cold event in the Holocene, it is very similar to these oscillations. The Eemian oscillation was noted also in the Bispingen profile in northern Germany, ~6500 years after the interglacial forestation. However, the course of vegetation changes there differs from that in Beckentin. The analysis of macroremains and Cladocera confirms the disappearance of rush and water vegetation in the water reservoir at that time in Beckentin. These results confirm that numerous climatic oscillations could have occurred in the Eemian Interglacial.

The floor: a voice of human lifeways—a geo-ethnographical study of historical and recent floors at Dolní Němčí Mill, Czech Republic

Is there a story that floor layers tell us about human history? A set of former floors were investigated using a combination of micromorphology, ethnography and macroremain analyses at a former mill in Dolni Němci in the eastern part of the Czech Republic. The floor layers had been accumulating since the end of the eighteenth century. A recently made earth floor, maintained for nearly 20 years, four times per year, in accordance with local knowledge passed down from mothers to daughters, formed the topmost layer. The main aim of the paper is: (1) to examine and contrast morphological characteristics of floors against known practices, which will be a useful reference for archaeological examples; (2) to compare the observations/findings to established knowledge of processes of floor formation. The structure of former floor layers preserved below the recent floor provided an indication of how floor care practices changed over the centuries, how these differed in different parts of the house and possible accumulation rate of the floor layer. This research illustrates that the micromorphological study of recent floors combined with ethnographic knowledge and macroremain analyses is a valuable methodological approach for the interpretation of archaeological case studies.

Plant Identification and Significance in Funeral Traditions Exemplified by Pillow Filling from a Child Crypt Burial in Byszewo (18th/19th centuries)

Plants have always played an important role in funeral customs. To understand their true meaning, close cooperation between the archaeologist and the archaeobotanist is needed, not only during the final interpretation, but from the very beginning, at the stage of collecting materials. In the article, plants’ identification, using both pollen and macroremains analysis, was described, based on one of the children’s burial from the Holy Trinity Church in Byszewo (18th/19th centu-ries). The filling of the coffin pillow consisted of numerous hop (Humulus lupulus) macroremains, the representation of which was very low in pollen sample. This is due to the fact that only female specimens of hop were inserted into the coffin. To determine the reason for using hops in funeral practices in Byszewo, ethnobotanical data was used. The following re-search indicates the need for the cooperation between two methods of plant identification. It will allow misinterpretations of botanical findings to be avoided.

Early Middle Ages Houses of Gien (France) from the Inside: Geoarchaeology and Archaeobotany of 9th–11th c. Floors

ABSTRACTAt Gien (France), indoor floors from early Middle Ages occupation (8th–10th c. AD) are very well preserved, providing a new reference for archaeological investigation in northern France. This site is located on an outcrop, 20 m above the Loire valley, where a 15th c. castle stands now. The medieval occupation combines high-status houses with crafting and agricultural areas. They constitute a new urban nucleus, which grew 2 km east from an ancient Roman settlement. During the rescue excavation, four buildings of different status were sampled and studied using an integrated approach, combining stratigraphy, micromorphology, chemical, macro-remain and phytolith analyses. Micromorphological investigations helped to identify 74 built floors, from 0.5 to 150 mm thick, made with transformed local clay or imported silty earth. Mineral floors were covered by vegetal ones, consisting of crop processing refuse. These litters include an abundance of phytoliths and some seeds, both produced by cultivated cereals, which were processed in situ, such as Triticum durum, Secale cereale and Hordeum vulgare. The refuse above the mineral and vegetal floors were trampled. They were produced not only by domestic activities, such as cooking and eating, but also by metallurgic activities and animal husbandry. The investigation of a contemporary pit indicated that, despite the large amount of refuse, floors were well maintained and regularly rebuilt. The spatial distribution of waste indicated that a single space could be dedicated to several activities, which were not necessarily separated by new floors. Moreover, the total absence of bioturbation allowed the study of a stage of dark earth formation, by comparing it to the contemporaneous mechanical disturbance of a part of the strata which occurred when building new floors. All these results give new evidence of the richness and the complexity of the early Middle Ages town, in addition to help identifying the activities which could take place in early castral areas.

The shining piece of the puzzle: evidence of plant use in the Late Palaeolithic

The Late Glacial site Lubrza 10 yielded new archaeological and use-wear data for discussing the development of plant-based technologies long before the occupation of the European Lowlands by Neolithic societies. More than 4000 Federmesser and Swiderian lithic artefacts were collected from the site, which is located on sandy kames adjacent to former water bodies. Use-wear analysis showed that abandoned tools were engaged in the activities well recognised as Palaeolithic ones, such as hunting, working hide, bone and other soft and hard materials. Microscopic observations have also produced some of the earliest evidence of processing non-woody plants in the North European Plain. There is a considerable number of artefacts with plant-like polish. Tools were used for whittling, splitting, scraping or stripping plant materials—motions that could have also been made while extracting of fibres and other raw materials necessary for making crafts. Morphological parameters of tools suggest a selection of flint implements, and their distribution shows the locations of processing plants. Results of previously conducted pollen and macro-remains analysis demonstrate the presence of a great variety of plants, including high silica content herbaceous species that covered the nearest vicinity of the site both during the Allerod and the Younger Dryas.

ANTHRACOLOGICAL EVALUATION AT ARCHAEOLOGICAL SITES OF GUARANI OCCUPATION - TAPHONOMIC COMPARISONS TO CONSTRUCT THE ENVIRONMENTAL HISTORY OF THE FORQUETA RIVER BASIN, RS, BRAZIL

Anthracological studies aim at the reconstruction of the paleoenvironment through the analysis and interpretation of macroremains of carbonized woods found in soils and archaeological sites that are related to previous human activities. The objective of this study was to compare the anthrocological and taphonomic data of the Archaeological Sites RS-T-101 and RS-T-114, located in the municipality of Marques de Souza, associated with pre-colonial occupancies of Guarani origin (archaeological sites RS-T-114 and RS-T-101, respectively, in 1,410-431 BP years and 1,411 -295 years BP, to build the environmental history of the Forqueta River basin based on taphonomic comparisons between these two sites. The traces analyzed were collected following methodology scaling and blasting, being conducted by the Archeology Section of the team’s Museum of Natural Sciences of the UNIVATES University Center. In this work, the images of carbon obtained under SEM were analyzed from both sites, which were the object of previous anthrachological studies, being used to perform the taphonomic comparisons. From the images, it can be inferred that the recovered charcoals in the sites had their anatomical characteristics well preserved, being possible to observe homogenization of the cellular walls, as well as the thickness of the walls demonstrates fires with temperatures that did not exceed 340ºC. Moreover, with this study it was possible to infer that the fragments found in these sites may have been originated by fires of anthropogenic origin because they are dispersed in the sedimentary matrix as well as in places with the presence of vestiges of fires.

Different parts of the same plants. Charcoals and seeds from Cova de les Cendres (Alicante, Spain)

The analysis of archaeobotanical macroremains from Cova de les Cendres (Teulada-Moraira, Alicante, Spain) allows a better understanding of the dynamic of the vegetation during the Upper Palaeolithic in the region. The anthracological sequence shows that Pinus type nigra/sylvestris dominates in the area during the Upper Palaeolithic, while more open formations of Juniperus and Fabaceae spread in Upper Solutrean and Early-Middle Magdalenian. The carpological analysis that has been carried out in the Middle Magdalenian level has brought to light different species of Juniperus (J. sabina, J. communis and J. oxycedrus). This information indicates that during the period, Cova de les Cendres had similar bioclimatic conditions to those currently prevailing in the Iberian summits of the supramediterranean belt. New 14C dates obtained from Juniperus sabina and Buglossoides cf. arvensis seeds as well as from charcoal fragments of Ephedra sp., presented in this paper, prove the existence of steppe landscapes during the Late Glacial.

Landscape transformations at the dawn of agriculture in southern Syria (10.7–9.9 ka cal. BP): Plant-specific responses to the impact of human activities and climate change

In southwest Asia, the accelerated impact of human activities on the landscape has often been linked to the development of fully agricultural societies during the middle and late Pre-Pottery Neolithic B (PPNB) period (around 10.2–7.9 ka cal. BP). This work contributes to the debate on the environmental impact of the so-called Neolitisation process by identifying the climatic and anthropogenic factors that contributed to change local and regional vegetation at the time when domesticated plants appeared and developed in southern Syria (around 10.7–9.9 ka cal. BP). In this work a multidisciplinary analysis of plant microremains (pollen and phytoliths) and macroremains (wood charcoal) is carried out along with stable carbon isotope discrimination of wood charcoals in an early PPNB site (Tell Qarassa North, west of the Jabal al-Arab area). Prior to 10.5 ka cal. BP, the results indicate a dynamic equilibrium in the local and regional vegetation, which comprised woodland-steppe, Mediterranean evergreen oak-woodlands, wetland vegetation and coniferous forests. Around 10.5–9.9 ka cal. BP, the elements that regulated the vegetation system changed, resulting in reduced proportions of arboreal cover and the spread of cold-tolerant and wetlands species. Our data show that reinforcing interaction between the elements of the anthropogenic (e.g. herding, fire-related activities) and climatic systems (e.g. temperature, rainfall) contributed to the transformation of early Holocene vegetation during the emergence of fully agricultural societies in southern Syria.

Algade 9 i Aalborg – Husenes udvikling fra vikingetid til nutid

Algade 9 i Aalborg – Husenes udvikling fra vikingetid til nutid

Palaeoecological reconstructions of vegetation successions in two contrasting former turbaries in the Netherlands and implications for conservation

Terrestrialization stages of mire vegetation are important to Dutch nature conservancy because of high biodiversity, particularly of base-rich fens. In recent turbaries, terrestrialization has however not shown development of such a stage, while existing rich fens show accelerated development towards species-poor Sphagnum-dominated vegetation, due to acidification and lack of base-rich water input. We apply the analysis of microfossils and macroremains in two peat cores in order to get a better understanding of terrestrialization in the past, and thus provide information on actual vegetation successions, which is directly relevant to nature conservation.The sampling site Stobbenribben showed a vegetation succession under relatively nutrient-poor conditions. The site at Westbroek underwent two peat digging events with one vegetation sequence starting in the 16th/17th century under nutrient-poor conditions, and another during the last century under relatively nutrient-rich conditions. Results from Westbroek showed that if the pond is shallow, rich-fen bryophytes develop relatively close to the bottom of newly formed peat, due to structural support by species such as Equisetum fluviatile and Schoenoplectus lacustris. For Stobbenribben, where the pond is much deeper, rich-fen bryophytes only established after development of a floating root mat with Stratiotes aloides. For both Stobbenribben and 20th century Westbroek, the base-rich fen stage took 60years to establish and remained for only 30years without additional input of base-rich water.Our palaeoecological records can be used to provide a longer temporal perspective to address conservation issues of base-rich fens. We conclude that digging up new turf ponds may not be enough to meet the obligations of the Natura 2000 Habitat Directive, if extra measures to prolong the lifespan of rich fens through e.g. temporary inundations with base-rich and nutrient-poor water are not taken into account.

The interpretation of pollen assemblages from medieval and post-medieval cesspits: New results from northern Belgium

Pollen has been studied from cesspits and other archaeological structures containing human faecal remains from different sites in northern Belgium, dating from the 12th to the 17th century. The results show that a large amount of information can be obtained by analysing pollen from such archaeological features. In addition, it is demonstrated that this information is largely complementary to the results obtained by the analysis of botanical macro-remains. The consumption of different food plants like herbs, spices and leafy vegetables, but also the use of honey, pharmaceutical plant use, and the use of peat fuel is reflected in the pollen assemblages from the studied archaeological structures. Most of this information cannot be obtained through the analysis of botanical macro-remains.

Last Glacial palaeoenvironments at Lascaux, southwest France, with special emphasis on MIS 4 (Ognon II interstadial)

A deep sequence of Last Glacial peats and slope deposits at the foot of the Lascaux hill was investigated using a large array of methods (geology, geophysics, 14C and OSL dating, palynology, macro-remain analysis, entomology). The peaty levels, OSL dated to between ~73 and 60ka, are found within a doline developed in a fluvial terrace of the Vézère River. These gyttja and chalky gyttja deposits covered by fen peat yielded abundant beetle and plant (seeds, pollen) remains. Palaeoecological reconstructions suggest a cool steppe with willow stands compatible with an interstadial of marine isotopic stage (MIS) 4, probably Ognon II (GI-19.2). Aquatic and hygrophilous plants (sedges), which reflect local vegetation, and their dependent insects are equally well represented in the record. The peats are covered by a sequence of slope deposits: a coarse-grained lower unit deposited during late MIS 4, overlain by a heavily decalcified sandy unit with a boreal soil complex correlated with early MIS 3, and finally, a coarse-grained, calcareous unit formed during late MIS 3 and MIS 2 in connection with slope dynamics typical of periglacial environments. The palaeosols yielded a small Middle Palaeolithic lithic assemblage.

Holocen history of vegetation at “Uroczysko Mokradła” (SW Poland) – paleobotanical research

Two peat bogs were studied in the Bory Dolnośląskie, the forest complex in Lower Silesia (Poland). An Instorf drill was used to collect two peat profiles from the deepest places. The macroremains analysis showed that after the initiation of peat-forming processes phytocoenoses responsible for the deposition of transition sphagnum peat were developed at both locations. Later on, the development of both peat bogs differed. The smaller peat bog continued to develop, whereas the big bog was shifted to ombrotrophic water regime. Therefore, phytocoenoses accumulated 1.3 m of <em>Sphagnum</em> peat. The peat-forming process was initiated at different times in both sites. For the smaller peat bog, it took place during the Atlantic period, while in the case of the larger peat bog – several thousand years later. The first identified forest phytocoenoses in the Atlantic period are mesophilic multi-species deciduous forests. Dry coniferous forests and mixed birch-pine forests grew in dry habitats. Riparian forests occupied lower grounds. In the Subboreal period, the oak–hazel communities initially developed and mixed coniferous forests were partially replaced by light oak forests. The encroachment of spruce, fir, hornbeam, and beech resulted in the development of dry ground forests, including beech–fir woods. The importance of riparian forests increased, whereas in dry grounds pine and mixed coniferous forests continued to occur. In the Subatlantic period, the transformations in forest communities were associated with the spread of hornbeam, beech, and fir and thereby vast fertile habitats were colonized by dry ground communities and beech woods. Pine and mixed forests as well as riparian forests were of lesser importance. Pollen records from the last 500 years showed the clear presence of humans. It was evident from the presence of cereal and weed pollen and from the disturbances in the pollen records caused by peat extraction in the Middle Ages.

Vegetation of the Ferdynandovian interglacial (MIS 13–15) based on plant macrofossils from a new profile of the stratotype site

Abstract Early Middle Pleistocene palaeolacustrine sediments of the Ferdynandów site (E Poland), serving as a stratotype for the Ferdynandovian interglacial, were subjected to a new drilling in 2011. The obtained profile, covering the late Sanian 1 glaciation, two interglacial successions (Ferdynandovian 1 and 2), the cold Ferdynandovian 1/2 interval, and the early Sanian 2 glaciation, permitted a high-resolution plant macroremains analysis, correlated with the results of a palynological examination. In detailed studies of plant macroremains from the profile, new taxa were discovered for the Polish Pleistocene flora: specifically, species that are extinct or not found nowadays in Poland. On the basis of the taxonomic diversity and occurrence of bioindicators, the division into warm and cold units of the Ferdynandovian interglacial as well as units associated with the Sanian 1 and 2 glaciations, already applied in pollen studies, were confirmed, and the periods were described in terms of plant macroremains. The Ferdynandovian succession is correlated with the Cromerian Complex (Cromerian III and IV) in the Early Middle Pleistocene of Western Europe, as well as with Marine Isotope Stages (MIS) 13-15. New geological and palaeobotanical data enabled a description of the palaeogeographic context and conditions of functioning, as well as the evolution of the interglacial lake at the stratotype site. The outcome of plant macroremains analysis is presented and interpreted in comparison with corresponding results previously obtained for the Ferdynandów B profile, studied by Janczyk-Kopikowa (1975). Particular consideration is given to similarities between the succession of Ferdynandów 2011 and the profiles of Ferdynandów B and the nearby drilled Łuków 3A. Study of the macrofossil flora of Ferdynandów 2011 revealed the presence of species absent in the archival Ferdynandów B. Taxa not formerly recorded in early Middle Pleistocene floras, such as Brasenia borysthenica, Aldrovanda borysthenica, Pilularia borysthenica, Caulinia goretskyi, and Potamogeton saryanensis, were identified. These species and other taxa of stratigraphic or climatic significance provided the basis for a more detailed description of the Ferdynandovian flora, afterwards compared with other stratigraphically corresponding Polish and European early Middle Pleistocene floras. The composition of characteristic taxa also was used for comparisons between two bimodal interglacial sequences, namely the Ferdynandovian (correlated with MIS 13-15) and Augustovian (correlated with MIS 19-21), and the Domuratovian succession (correlated with MIS 17).

Palaeoclimatic changes in the Holsteinian Interglacial (Middle Pleistocene) on the basis of indicator-species method – Palynological and macrofossils remains from Nowiny Żukowskie site (SE Poland)

Analysis of pollen and macroremains from the Mazovian/Holsteinian interglacial in the Nowiny Żukowskie succession enabled the identification of numerous climatic indicators. The applied indicator-species method allowed for a detailed description of climatic changes, i.e. warmings and coolings that occurred in this interglacial. The determined ranges of temperatures that prevailed in particular periods provide a basis for an assumption that it was probably the warmest interglacial of the Pleistocene. The estimated temperature of the warmest month during the climatic optimum (Abies–Carpinus period) may have been 2–4 C° higher than in the warmest period of the Holocene. Moreover, numerous changes in humidity, evidenced by frequent oscillations from oceanic to continental climate, are particularly important. The cool periods are generally marked by the presence of Betula nana, Selaginella selaginoides and S. helvetica, heliophytes such as Helianthemum nummularium, and shrubs of Ephedra and Hippophaë rhamnoides. The warmest period is documented by the occurrence of thermophylous trees, e.g. Pterocarya fraxinifolia, Juglans, shrubs, e.g. Buxus sempervirens and Vitis sylvestris, as well as Hedera helix, Viscum, and aquatic plants, e.g. Brasenia borysthenica.