ABSTRACT Changes in global climate from the Late Pennsylvanian to early Permian resulting from the destabilization of ice sheets in the Southern Hemisphere had important effects on terrestrial landscapes around the Paleozoic world. In the northern Appalachian Basin (U.S.A.), evidence for this transition and its effects on terrestrial systems is preserved in numerous paleosols in the Conemaugh, Monongahela, and Dunkard groups that formed in various alluvial environments. Several studies along a 50 km west–east and 40 km north–south transect through southeast Ohio and southwest West Virginia resulted in the recognition of 24 different pedotypes. An up-section shift in pedotypes from Argillisols to Vertisols and Calcisols as well as an overall upward increase in the diversity of pedotypes records a significant change in soil-forming processes and landscapes. The landscapes of the Late Pennsylvanian were largely characterized by well-developed, clay-rich Alfisols of woodland ecosystems on distal floodplains bordered by poorly developed Inceptisols of early successional or marshland ecosystems on proximal floodplains. By the early Permian, these landscapes began to experience strongly seasonal climates and increasingly prolonged dry periods, resulting in the production of pronounced shrink–swell features and well-developed carbonate horizons. These landscapes were characterized by Vertisols and Inceptisols of highly heterogeneous brakeland ecosystems, with rapid changes in sedimentation, hydrology, and vegetation over short distances. Climatic conditions were prone to fluctuations between wet and dry states during the transition, resulting in a highly heterogeneous landscape that changed significantly through time.