Acquisition Of Characters Research Articles

New specimens of Bunaia woodwardi Clarke, 1919 (Euchelicerata): a new member of Offacolidae providing insight supporting the Arachnomorpha.

The rapid early diversification of arthropods has made understanding internal relationships within the group fiendish. Particularly unresolved is the origin of Euchelicerata, a clade consisting of the Prosomapoda (comprising the extant Xiphosura and Arachnida and the extinct Chasmataspidida, Eurypterida and synziphosurines) and the extinct Offacolidae. Here we describe new material of the Silurian 'synziphosurine' Bunaia woodwardi Clarke, 1919 that reveals previously unknown features of its ventral anatomy: a pair of elongated chelicerae in the prosoma, followed posteriorly by five pairs of biramous appendages, a first pre-abdomen somite bearing a pair of paddle-like uniramous appendages (exopods) and a ventral pretelsonic process. Phylogenetic analyses retrieve B. woodwardi as a member of Offacolidae closely related to Setapedites abundantis from the Early Ordovician Fezouata Biota. An anatomical comparison of the pretelsonic process of B. woodwardi, also present in Setapedites, with the posterior trunk morphologies of other Offacolidae, Habeliida and Vicissicaudata, suggests a possible homologous appendicular origin. This proposed apomorphic character supports a monophyletic Arachnomorpha, formed of Vicissicaudata, Habeliida and Euchelicerata. The establishment of this new homology could help to clarify the highly enigmatic phylogeny at the base of the euchelicerates as well as the sequence of character acquisition during their early evolution.

Design and practice of human-machine cooperative international Chinese character teaching in ChatGPT application

The incorporation of artificial intelligence (AI) into language education has created new opportunities for improving the instruction and acquisition of Chinese characters. Nevertheless, the cognitive difficulties linked to the acquisition of Chinese characters, such as their intricate visual features and lack of clear meaning, necessitate thoughtful deliberation when developing AI-supported learning interventions. The objective of this project is to explore the capacity of a collaborative method between humans and machines in teaching Chinese characters, utilising the advantages of both human expertise and AI technology. We specifically investigate the utilisation of ChatGPT, a substantial language model, for the creation of instructional materials and evaluation methods aimed at teaching Chinese characters to individuals who are not native speakers. The study utilises a mixed-methods approach, which involves both qualitative examination of lesson plans created by ChatGPT and quantitative evaluation of student learning outcomes. The results indicate that the suggested framework for human-machine collaboration can successfully tackle the cognitive difficulties associated with learning Chinese characters, resulting in enhanced learner involvement and performance. Nevertheless, the research also emphasises the constraints of AI-generated material and the significance of human involvement in guaranteeing the accuracy and dependability of educational interventions. This research adds to the expanding collection of literature on AI-assisted language learning and offers practical insights for educators and instructional designers who aim to use AI tools into Chinese language curriculum. The results emphasise the necessity of employing a multi-disciplinary strategy in AI-supported language learning, incorporating knowledge from cognitive psychology, educational technology, and second language acquisition.

Organ systems of a Cambrian euarthropod larva

The Cambrian radiation of euarthropods can be attributed to an adaptable body plan. Sophisticated brains and specialized feeding appendages, which are elaborations of serially repeated organ systems and jointed appendages, underpin the dominance of Euarthropoda in a broad suite of ecological settings. The origin of the euarthropod body plan from a grade of vermiform taxa with hydrostatic lobopodous appendages (‘lobopodian worms’)1,2 is founded on data from Burgess Shale-type fossils. However, the compaction associated with such preservation obscures internal anatomy3–6. Phosphatized microfossils provide a complementary three-dimensional perspective on early crown group euarthropods7, but few lobopodians8,9. Here we describe the internal and external anatomy of a three-dimensionally preserved euarthropod larva with lobopods, midgut glands and a sophisticated head. The architecture of the nervous system informs the early configuration of the euarthropod brain and its associated appendages and sensory organs, clarifying homologies across Panarthropoda. The deep evolutionary position of Youti yuanshi gen. et sp. nov. informs the sequence of character acquisition during arthropod evolution, demonstrating a deep origin of sophisticated haemolymph circulatory systems, and illuminating the internal anatomical changes that propelled the rise and diversification of this enduringly successful group.

A Fossil Record of Spores before Sporophytes

Because their resistant, sporopolleninous walls preserve a record of morphogenetic change during spore formation, fossil cryptospores provide a direct physical record of the evolution of sporogenesis during the algal–plant transition. That transition itself is a story of the evolution of development—it is not about phylogeny. Here, we review the fossil record of terrestrially derived spore/cryptospore assemblages and attempt to place these microfossils in their evolutionary context with respect to the origin of complex multicellularity in plants. Cambrian cryptospores show features related to karyokinesis seen in extant charophytes, but they also possess ultrastructure similar to that seen in liverworts today. Dyadospora, a cryptospore dyad recovered from sporangia of Devonian embryophytes, first occurs in the earliest Ordovician. Tetrahedraletes, a likely precursor to the trilete spore, first occurs in the Middle Ordovician. These fossils correspond to evolutionary novelties that were acquired during a period of genome assembly prior to the existence of upright, axial sporophytes. The cryptospore/spore fossil record provides a temporal scaffold for the acquisition of novel characters relating to the evolution of plant sporogenesis during the Cambrian–Silurian interval.

The inner ear and stapes of the basal mammaliaform Morganucodon revisited: new information on labyrinth morphology and promontorial vascularization

Abstract Based on high-resolution computed tomography scanning, we provide new insights into the inner ear and stapedial morphology of Morganucodon from the Early Jurassic of St Brides. At the base of mammaliaforms, Morganucodon plays a pivotal role in understanding the sequence of character acquisition from basal cynodonts to mammals, including the detachment of the middle ear and the evolution of high-frequency hearing. Advancements in imaging technology enabled us to revise or newly describe crucial anatomy that was not accessible for the original description of Morganucodon. Based on 37 petrosals, we can confirm that the apex of the cochlear canal is expanded in Morganucodon, suggestive of a lagena macula. A gently raised crest along the abneural margin is reminiscent of (although much shallower than) the secondary lamina base of other Mesozoic mammaliaforms. The venous circum-promontorial plexus, which surrounded the inner ear in several basal mammaliaforms, was connected to the cochlear labyrinth in Morganucodon through numerous openings along the secondary lamina base. Two petrosals contain fragmentary stapes, which differ substantially from previously described isolated stapes attributed to Morganucodon in having peripherally placed crura and an oval and bullate footplate. Based on the revised stapedial morphology, we question the traditional view of an asymmetrical bicrural stapes as the plesiomorphic condition for Mammaliaformes.

New records of early Paleocene (earliest Torrejonian) plesiadapiforms from northeastern Montana, USA, provide a window into the diversification of stem primates

Plesiadapiforms (putative stem primates) appear in the fossil record shortly after the Cretaceous/Paleogene boundary and subsequently radiated throughout the Paleocene into a taxonomically and ecomorphologically diverse group. The oldest known plesiadapiforms come from early Puercan (the oldest North American Land Mammal ‘age’ [NALMA] of the Cenozoic) deposits in northeastern Montana, and all records of Puercan plesiadapiforms are taxonomically restricted to members of the Purgatoriidae and the enigmatic genus Pandemonium. Plesiadapiform diversity substantially increased in the following Torrejonian NALMA, but the sparse record of faunas between the Puercan and the well-known middle and late Torrejonian has hampered our understanding of this important interval in early primate evolution. Here we report new plesiadapiform dental fossils from early Torrejonian (To1) deposits from the Tullock Member of the Fort Union Formation in northeastern Montana that record several poorly known taxa including members of the Purgatoriidae, Paromomyidae and Pandemonium, and that document the largest and most diverse assemblage of To1 plesiadapiforms known. We describe a new species of the purgatoriid Ursolestes (Ursolestes blissorum, sp. nov.) that represents the largest plesiadapiform known from the early Paleocene and, among other taxa, provides additional evidence that the temporal range of purgatoriids extended into the Torrejonian. Large sample sizes of the oldest known paromomyid, Paromomys farrandi, allowed us to document intraspecific variability and one undescribed tooth locus. Our observations illuminate changes in dental morphology of some taxa that occurred in To1 and may inform the acquisition of certain diagnostic plesiadapiform dental characters. We evaluate plesiadapiform species richness, mean body mass and body-mass disparity through the Paleocene and reveal unrecognized levels of richness in To1 and a general trend of stable body mass and body-mass disparity. Our findings contribute to documented patterns of plesiadapiform provincialism in the early Paleocene and shed light on the early stages of their Torrejonian radiation.

Using AR-assisted approach in Chinese character acquisition for adult CFL learners

Using AR-assisted approach in Chinese character acquisition for adult CFL learners

Study on Chinese Character Acquisition Strategies for Beginner Level Second Language Learners

More and more scholars are paying attention to the acquisition strategies of Chinese as a second language. This is because when teachers fully understand learners' acquisition strategies, they can use this as an entry point to formulate teaching methods suitable for learners, and the teaching effect will naturally get twice the result with half the effort. For beginners, Chinese characters are considered to be the most difficult part of learning Chinese as a second language. Of course, it is necessary to study their acquisition strategies in detail. Combined with the characteristics of primary second language learners' Chinese character acquisition strategies, teachers can improve the shortcomings of Chinese character teaching design and implement more efficient and targeted classroom teaching.

Origin and Early Evolution of Echinoderms

Echinoderms are a major group (phylum) of invertebrate animals with a rich fossil record stretching back to the Cambrian period, approximately 518 million years ago. While all modern species are characterized by pentaradial (i.e., fivefold) symmetry, Cambrian echinoderms also include taxa with different types of symmetry (e.g., bilateral symmetry). These distinct forms were present from very early in the phylum's history, demonstrating that the initial diversification of echinoderm body plans was extremely rapid. The phylogenetic relationships of Cambrian echinoderms have long been debated, hindering efforts to reconstruct the evolution of the phylum, but recent analyses have consistently recovered bilaterally symmetrical forms as the earliest-diverging echinoderms. This reveals the sequence of character acquisition in echinoderm evolution, indicating that radial symmetry is a derived character of the group, which evolved after the acquisition of a mineralized skeleton. Cambrian echinoderms were adapted to diverse modes of life, with ecology an important factor shaping their early evolution. However, the reasons why echinoderms evolved their unique pentaradial body plan remain unclear. ▪The Cambrian fossil record provides valuable insights into the origin and early evolution of echinoderms over half a billion years ago.▪Cambrian echinoderms were morphologically diverse, with several extinct groups exhibiting character combinations that distinguish them from living species.▪Phylogenetic analyses of bilateral, asymmetrical, triradial, and pentaradial fossils have allowed us to decipher the assembly of the modern echinoderm body plan.▪Echinoderms became ecologically diverse early in their history, with varied modes of feeding, locomotion, and attachment.

The effectiveness of visual instructions in the memorization of Chinese Semantic Radicals by Beginners

Abstract The acquisition of Chinese Characters (CCs) is challenging for learners. Chinese Semantic Radicals (SRs) are semantic components of CCs. The memorization of SRs facilitates the acquisition of CCs when learning Chinese as a second language. This study investigates the effectiveness of Visual Instructions (VIs) in the memorization of SRs by beginners. A preliminary study and three other studies have been conducted. The results have shown that the use of color-coded illustrations helps learners remember the meanings, shapes, and positions of SRs. This could be seen in the significantly increased accuracy of the responses in memory tests. These findings are an important contribution to research on VIs and on the memorization of SRs by beginners.

Chinese heritage children learning to read characters: The effects of group reading and frequency of character exposure

Learning to read Chinese has been the main focus of Chinese heritage language education. The frequency of exposure to orthography–phonology and orthography–semantics correspondences and radical awareness play a significant role in character learning. However, limited exposure to characters in Chinese textbooks and the lack of explicit instruction on phonetic radicals have been proposed as limitations of conventional instruction methods used with Chinese heritage language learners. The current study assessed the efficacy of providing frequent character exposure in group reading activities for Chinese heritage children in their early learning to read Mandarin Chinese through a teaching method called Progressive Character Reading. The effectiveness of the teaching method on Chinese character acquisition and Chinese reading comprehension was examined through a quasi-experimental framework that compared a group of children undergoing the new teaching method (experimental group, n = 20) with a group of learners experiencing the traditional textbook-based instruction (control group, n = 20). Significant differences in gains between these two groups were found in measures of Chinese character reading, character–picture matching and Chinese sentence comprehension. These results suggest that group reading activities with frequent character exposure can increase learning for Chinese heritage language learners over that associated with conventional textbook-based instruction.

Foreword

Dear Linguistic Enthusiasts and Scholars, It is with great pleasure that we present this year’s summer issue of the scientific journal of Acta Linguistica Asiatica. With it, we embark on a journey to explore the rich and diverse linguistic tapestry of the Japanese, Chinese, and Korean languages, delving into their contemporary, pedagogical, historical, and other significance. We extend our heartfelt gratitude to the authors, reviewers, and editorial team whose unwavering commitment has made this journal possible. It is our hope that the ALA journal will continue to serve as a platform to foster intellectual exchange and contribute to the advancement of linguistic knowledge in this dynamic and ever-evolving field. This issue offers 4 research articles and one book review. It opens with the article “Distant Co-occurrence Patterns of Connectives: a Corpus Study of Formulaicity in Japanese” written by Andrej BEKEŠ, Bor HODOŠČEK, NISHINA Kikuko, and ABEKAWA Takeshi. In the article, the authors examined two-item and multi-item distant co-occurrence patterns of connectives in written Japanese using corpus research methods. The analysis shows these patterns contribute to discourse development and prediction, resembling Bourdieu’s habitus. In the following article “Teaching Both Simplified and Traditional Characters to Learners of Chinese as L2”, LI Xiao examined Chinese character acquisition in second language learning. It discusses different approaches - prioritizing simplified or traditional characters, teaching both together, or delaying character learning. Results show that most learners prefer simplified characters, some equally acquired both, while fewer prefer traditional characters. Text comprehension is better with characters than with Hanyu Pinyin. The article “Exclamation in Late Archaic Chinese” by WANG Aiqing delved into exclamations in Late Archaic Chinese (LAC), which consist of sentence exclamations and exclamatives. Sentence exclamations assert propositions with or without surprise. The author proposes that exclamatives with degree adverbials he and heqi do not count as true exclamatives in LAC. The last research article in this issue, contributed by Klaudija LUKMAN and RYU Hyeonsook is entitled “Korean Honorifics in Flux: a Case Study of Seoul National University LEI Textbooks”. By examining Korean language textbooks (2000-2019), this research showed the link between society and politeness styles. Notable shifts reveal the replacement of formal hasipsioche with polite haeyoche, reflecting changes in Korean culture and society. Last but not least, Nina GOLOB reviewed the book “Uvod u znanost o japanskom jeziku: Osnovna obilježja, glasovni sustav i leksički slojevi.” Its primary contribution lies in its methodical exploration of diverse facets of the Japanese language and the accomplishments of linguistic research related to it, all while considering the perspective of Croatian and other Slavic languages. The book is ideal for Croatian-speaking Japanese students and linguistics enthusiasts. At this point, it is worth mentioning that in this era of rapid technological advancement, we wish to highlight the significant influence of Artificial Intelligence (AI) on linguistic studies. The emergence of AI has brought about a revolution in our methodologies, granting us the capability to analyze extensive linguistic data in real-time and fostering cross-linguistic collaborations with the aid of language translation tools. While we appreciate the immense potential that AI offers, we must tread carefully, striking a balance between leveraging AI’s potential and preserving the essence of human-driven analysis. Ethical considerations surrounding AI integration also deserve our attention. Editors and Editorial board wish the regular and new readers of the ALA journal a pleasant read full of inspiration, and a rise of new research ideas inspired by these papers. Editors

A Study of Chinese character acquisition in the digital age

A Study of Chinese character acquisition in the digital age

The early language gap between first- and second-language learners: acquisition of Chinese characters among preschoolers.

For culturally and linguistically diverse children, early second language (L2) development is important for school achievement and social inclusion. These children face challenges in acquiring L2, especially in Hong Kong, where the dominant Chinese language contrasts strongly with their home languages. Studies that compared the language abilities of first language (L1) and L2 students in English-speaking contexts have reported young L2 learners' disadvantage in using the dominant language in oral language and comprehension at school entry. The findings raise the question of whether L2 learners who fall behind their L1 peers in language abilities will be further disadvantaged, showing a weaker development gradient. This study used the Chinese Character Acquisition Assessment (CCAA) to compare character acquisition of 491 L2 children aged from 3 to 6 years against that of 240 of their L1 peers from Hong Kong kindergartens. The CCAA is comprised of six subtests and assesses children's abilities to make associations among character written form (orthography), sound, and meaning. Results showed that L2 learners had greater development in meaning and sound associations across class levels, implying that they may first develop oral language related abilities. In addition, results indicate that diverging gaps between L1 and L2 learners' Chinese character acquisition existed across class levels for the associations involving written character form, but not in regard to associations between character meaning and sound. This study highlights the Chinese learning needs of L2 preschoolers and provides understanding of their abilities in mapping among character written forms, sounds, and meanings. The findings suggest the importance of supporting L2 children's oral language at earlier stages of Chinese learning, and the need to provide instructional support to compensate for their relative weakness in literacy at school entry.

Plastome phylogenomics provide new perspective into the phylogeny and evolution of Betulaceae (Fagales)

BackgroundBetulaceae is a relatively small but morphologically diverse family, with many species having important economic and ecological values. Although plastome structure of Betulaceae has been reported sporadically, a comprehensive exploration for plastome evolution is still lacking. Besides, previous phylogenies had been constructed based on limited gene fragments, generating unrobust phylogenetic framework and hindering further studies on divergence ages, biogeography and character evolution. Here, 109 plastomes (sixteen newly assembled and 93 previously published) were subject to comparative genomic and phylogenomic analyses to reconstruct a robust phylogeny and trace the diversification history of Betulaceae.ResultsAll Betulaceae plastomes were highly conserved in genome size, gene order, and structure, although specific variations such as gene loss and IR boundary shifts were revealed. Ten divergent hotspots, including five coding regions (Pi > 0.02) and five noncoding regions (Pi > 0.035), were identified as candidate DNA barcodes for phylogenetic analysis and species delimitation. Phylogenomic analyses yielded high-resolution topology that supported reciprocal monophyly between Betula and Alnus within Betuloideae, and successive divergence of Corylus, Ostryopsis, and Carpinus-Ostrya within Coryloideae. Incomplete lineage sorting and hybridization may be responsible for the mutual paraphyly between Ostrya and Carpinus. Betulaceae ancestors originated from East Asia during the upper Cretaceous; dispersals and subsequent vicariance accompanied by historical environment changes contributed to its diversification and intercontinental disjunction. Ancestral state reconstruction indicated the acquisition of many taxonomic characters was actually the results of parallel or reversal evolution.ConclusionsOur research represents the most comprehensive taxon-sampled and plastome-level phylogenetic inference for Betulaceae to date. The results clearly document global patterns of plastome structural evolution, and established a well-supported phylogeny of Betulaceae. The robust phylogenetic framework not only provides new insights into the intergeneric relationships, but also contributes to a perspective on the diversification history and evolution of the family.

The developing bird pelvis passes through ancestral dinosaurian conditions.

Living birds (Aves) have bodies substantially modified from the ancestral reptilian condition. The avian pelvis in particular experienced major changes during the transition from early archosaurs to living birds1,2. This stepwise transformation is well documented by an excellent fossil record2-4; however, the ontogenetic alterations that underly it are less well understood. We used embryological imaging techniques to examine the morphogenesis of avian pelvic tissues in three dimensions, allowing direct comparison with the fossil record. Many ancestral dinosaurian features2 (for example, a forward-facing pubis, short ilium and pubic 'boot') are transiently present in the early morphogenesis of birds and arrive at their typical 'avian' form after transitioning through a prenatal developmental sequence that mirrors the phylogenetic sequence of character acquisition. We demonstrate quantitatively that avian pelvic ontogeny parallels the non-avian dinosaur-to-bird transition and provide evidence for phenotypic covariance within the pelvis that is conserved across Archosauria. The presence of ancestral states in avian embryos may stem from this conserved covariant relationship. In sum, our data provide evidence that the avian pelvis, whose early development has been little studied5-7, evolved through terminal addition-a mechanism8-10 whereby new apomorphic states are added to the end of a developmental sequence, resulting in expression8,11 of ancestral character states earlier in that sequence. The phenotypic integration we detected suggests a previously unrecognized mechanism for terminal addition and hints that retention of ancestral states in development is common during evolutionary transitions.

A three-eyed radiodont with fossilized neuroanatomy informs the origin of the arthropod head and segmentation

In addition to being among the most iconic and bizarre-looking Cambrian animals, radiodonts are a groupthat offers key insight into the acquisition of the arthropod body plan by virtue of their phylogenetic divergence prior to all living members of the phylum. Nonetheless, radiodont fossils are rare and often fragmentary, and contentions over their interpretation have hindered resolution of important evolutionary conundrums. Here, we describe 268 specimens of Stanleycaris hirpex from the Cambrian Burgess Shale, including many exceptionally preserved whole-body specimens, informing the most complete reconstruction of a radiodont to date. The trunk region of Stanleycaris has up to 17 segments plus two pairs of filiform caudal blades. The recognition of dorsal sclerotic segmentation of the trunk cuticle and putative unganglionated nerve cords provides new insight into the relative timing of acquisition of segmental traits, the epitome of the arthropod body plan. In addition to the pair of stalked lateral eyes, the short head unexpectedly bears a large median eye situated behind a preocular sclerite on an anteriorly projecting head lobe. Upon re-evaluation, similar median eyes can be identified in other Cambrian panarthropods demonstrating a deep evolutionary continuity. The exquisitely preserved brain of Stanleycaris is consistent with the hypothesized deutocerebral innervation of the frontal appendages, reconciling neuroanatomical evidence with external morphology in support of an ancestrally bipartite head and brain for arthropods. We propose that the integration of this bipartite head prior to the acquisition of most segmental characters exclusively in the arthropod trunk may help explain its developmental differentiation.

The origin and early evolution of arthropods.

The rise of arthropods is a decisive event in the history of life. Likely the first animals to have established themselves on land and in the air, arthropods have pervaded nearly all ecosystems and have become pillars of the planet's ecological networks. Forerunners of this saga, exceptionally well-preserved Palaeozoic fossils recently discovered or re-discovered using new approaches and techniques have elucidated the precocious appearance of extant lineages at the onset of the Cambrian explosion, and pointed to the critical role of the plankton and hard integuments in early arthropod diversification. The notion put forward at the beginning of the century that the acquisition of extant arthropod characters was stepwise and represented by the majority of Cambrian fossil taxa is being rewritten. Although some key traits leading to Euarthropoda are indeed well documented along a diversified phylogenetic stem, this stem led to several speciose and ecologically diverse radiations leaving descendants late into the Palaeozoic, and a large part, if not all of the Cambrian euarthropods can now be placed on either of the two extant lineages: Mandibulata and Chelicerata. These new observations and discoveries have altered our view on the nature and timing of the Cambrian explosion and clarified diagnostic characters at the origin of extant arthropods, but also raised new questions, especially with respect to cephalic plasticity. There is now strong evidence that early arthropods shared a homologous frontalmost appendage, coined here the cheira, which likely evolved into antennules and chelicerae, but other aspects, such as brain and labrum evolution, are still subject to active debate. The early evolution of panarthropods was generally driven by increased mastication and predation efficiency and sophistication, but a wealth of recent studies have also highlighted the prevalent role of suspension-feeding, for which early panarthropods developed their own adaptive feedback through both specialized appendages and the diversification of small, morphologically differentiated larvae. In a context of general integumental differentiation and hardening across Cambrian metazoans, arthrodization of body and limbs notably prompted two diverging strategies of basipod differentiation, which arguably became founding criteria in the divergence of total-groups Mandibulata and Chelicerata. The kinship of trilobites and their relatives remains a source of disagreement, but a recent topological solution, termed the 'deep split', could embed Artiopoda as sister taxa to chelicerates and constitute definitive support for Arachnomorpha. Although Cambrian fossils have been critical to all these findings, data of exceptional quality have also been accumulating from other Palaeozoic Konservat-Lagerstätten, and a better integration of this information promises a much more complete and elaborate picture of early arthropod evolution in the near future. From the broader perspective of a total-evidence approach to the understanding of life's history, and despite persisting systematic debates and new interpretative challenges, various advances based on palaeontological evidence open the prospect of finally using the full potential of the most diverse animal phylum to investigate macroevolutionary patterns and processes.

The closest evolutionary relatives of pterosaurs: What the morphospace occupation of different skeletal regions tell us about lagerpetids.

Exquisite discoveries and new interpretations regarding an enigmatic group of cursorial avemetatarsalians led to a new phylogenetic hypothesis regarding pterosaur affinities. Previously thought to be dinosaur precursors, lagerpetids are now considered to be the closest relatives to pterosaurs. This new hypothesis sheds light on a new explorable field, especially regarding the character acquisition and evolution within the pterosaur lineage. In the present study, the morphospace occupation of distinct skeletal regions of lagerpetids within the morphological spectrum of avemetatarsalians is investigated. This approach indicates which portions of the skeleton are more similar to the anatomy of pterosaurs and which portions present different homoplastic signals. The analyses demonstrate that the craniomandibular traits of lagerpetids are pterosaur-like, the pectoral girdle and forelimb are dinosauromorph-like and the axial skeleton and the pelvic girdle and hindlimb are unique and highly specialized among the analyzed sample. So, despite the close phylogenetic relationships, the postcranial skeleton of lagerpetids and pterosaurs are very different. The occurrence of two distinct and highly specialized groups of pterosauromorphs coexisting with a wide ecological range of dinosauromorphs during the Triassic suggests pressure for new niches occupation.

The affective embeddings of gacha games: Aesthetic assemblages and the mediated expression of the self

This article argues that the motivations for investing money in gacha games can be a function of the affective embedding of players within the game, and the game within broader circuits of cultural affinity and appeal. While research on gacha games – and the specific role of loot boxes therein – has emphasised their associations with gambling, I contend that affect is another trigger that can motivate seemingly irrational playing behaviours. The affective embeddings of gacha games motivate players to curate aesthetic assemblages of virtual content that enable the mediated expression of the self. Drawing on qualitative data generated among young Singapore-based players of gacha games, I explore how the acquisition of characters, skins and collections can be motivated by the emotional payoff that comes from relationality rather than gambling.