Summary. The African Scytopetalaceae, hitherto described as exstipulate, do in fact produce minute, earlycaducous stipules. This supports the inclusion of the stipulate American genus Asteranthos in Scytopetalaceae. The character 'presence or absence of stipules' plays an important role in the identification and classification of flowering plants. Their presence or absence is usually constant for a family as a whole, but in some families (e.g. Flacourtiaceae, Sapindaceae, Sapotaceae) it is valid at a lower level, i.e., constant for genera or for species within genera. In some taxa stipules are very small and/or early-caducous, leaving hardly any trace of their previous presence. Such taxa may easily be taken to be exstipulate. Scytopetalaceae is a small African family of 19 species in 5 genera: Brazzeia Baill., Oubanguia Baill., and Scytopetalum Pierre ex Engl. each with 3 species, Rhaptopetalum Oliv. with 9 and Pierrina Engl. with a single species. The family is best represented in the Cameroon - Gabon area where all the 5 genera, and 16 species, occur (Letouzey 1978a, b), followed by the area of the Flore d'Afrique Centrale where 7 species in 4 genera are found (Germain 1963), and West Tropical Africa where there are 5 species, also in 4 genera (Keay 1958). In Angola (Exell & Mendonqa 1951) and Uganda (Verdcourt 1968) only a single species is present. All treatments of the Scytopetalaceae for the African floras (Exell & Mendonqa 1951; Germain 1963; Keay 1958; Letouzey 1978a, b; Verdcourt 1968) describe the family as exstipulate. Appel (1996), who studied the morphology of this family in detail, also described it as such. When sorting sterile material from Gabon into families I found minute stipules in a specimen (Breteler et al. 14604) which was therefore first placed in Flacourtiaceae. When it could not be matched with any known species of this family, a closer look revealed a texture of the dried leaves that pointed towards Scytopetalaceae. In fact, it represented Scytopetalum klaineanum Pierre ex Engl. (Fig. 1). All species of this family were then investigated to see whether the presence of stipules was a character of the family. Stipules could not be found only in Oubanguia laurifolia (Pierre ex De Wild.) Tiegh. and in Rhaptopetalum belingense Letouzey, almost certainly because the material investigated was too mature to show these minute, fragile, early-caducous organs. Their presence can only be established on young immature shoots. In fact no distinguishable trace is left and it is understandable that the family has hitherto
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