What the pig ate: A microbotanical study of pig dental calculus from 10th–3rd millennium BC northern Mesopotamia

Abstract

One of the main questions that zooarcheologists have attempted to answer in their studies of ancient agropastoral economies relates to animal diet. Starch granules and phytoliths, which derive from the plant foods consumed over the course of an animal's life, become imbedded in dental calculus and thus offer direct clues about diet. In this paper, we investigate pig diet with an eye toward understanding husbandry strategies in northern Mesopotamia, the region in which pigs were first domesticated, from the Epipaleolithic though the Early Bronze Age. Our data reveal that pigs consumed an assortment of plant foods, including grasses, wild tubers, acorns, and domestic cereals. Although poor preservation plagued the identification of plant microremains at Epipaleolithic (10th millennium cal. BC) Hallan Çemi, the identification of a diet based on tubers and grasses matches models of wild boar diet. Pigs at 6th millennium Domuztepe, 5th millennium Ziyadeh, and 4th millennium Hacinebi consumed cereals, particularly oats (Avena sp.) and barley (Hordeum sp.), as well as wild plant food resources. Several of the cereal starch granules showed evidence of cooking, indicating that pigs had access to household refuse beginning at least in the late Neolithic. Moreover, calculus from morphologically wild specimens also contained cooked cereal grains. This points to a close relationship between wild boar populations and human settlements in the Neolithic and beyond. Preservation was poor for 3rd millennium sites in the study, including Atij, Raqa'i, Ziyadeh, and Leilan, but the available data suggest that pigs ate oats, barley and other Triticeae (the tribe that includes wheat, barley, and goatgrass), and other grasses. This may represent foddering practices in the Early Bronze Age.