Abstract
ABSTRACT Much of the visceral skeleton (jaws, hyoid arch, and branchial arches) and superficial orobranchial musculature is described from an unique three-dimensionally preserved cladoselachian from the Upper Devonian Chattanooga Shale of Tennessee. The mandibular joint lacks the complexity seen in modern chondrichthyans. The hyoid arch apparently helped support the mandibular Suspensorium, but may have had only a weak craniohyoidean attachment. Five branchial arches are present, each supporting elongate branchial rays that project beyond the primary hemibranch lamellae. Impressions of branchiomeric musculature conclusively demonstrate that there was no continuous gill opening between the jaws and hyoid arch; this cladoselachian is not aphetohyoidean.
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