Vipera berus (Linnaeus, 1758) remains from the Late Pleistocene of Slovakia

Abstract

Completely preserved specimens of fossil snakes are extremely rare and ophidian palaeontologists are usually dependent only on disarticulated elements of a postcranial skeleton. Here we present an unusually well-preserved specimen of a small viperid snake from the Late Pleistocene firm travertine at the famous Gánovce-Hrádok Neanderthal mound in Slovakia. The complex study of both cranial and axial skeleton with well-preserved maxilla and basiparasphenoid confirms the presence of a viper from theVipera berusspecies group, and recent distribution ofV. berusspecies complex members supports identification of these preserved remains as belonging to the common adder,V. berus(Linnaeus, 1758). Associated faunal assemblages of the MFG-C and D mammalian fauna groups reported from the firm travertine indicate a humid climate in a predominantly woodland environment with typical forest species in the Gánovce-Hrádok vicinity throughout the Eemian optimum, and mixed forest and steppe environments in the late Eemian to early Weichselian stages. Occurrence ofV. berusdocuments the presence of open or semi-open biotopes with low vegetation. AlthoughV. berusoccurs in the Quaternary glacial/interglacial cycle and throughout the entire warm part, it never dominated herpetofaunal assemblages during the climatic optimum. Therefore, the presence ofV. berusmost likely indicates late Eemian or early Weichselian (interstadial) climatic conditions.