Abstract
Six ovulate, permineralized cones, four cone-scale complexes, and one isolated seed are described from the Upper Cretaceous Upper Yezo and Miho Groups from Hokkaido, Japan, and Saghalien, Russia. Cones are spherical, 3.5-6.0 cm in diameter, with prominent thick bracts. Thick ovuliferous scales are almost completely united with the bracts and are not readily distinguishable on all scales. Cone-scale tissues consist of prominent abaxial and adaxial sclerenchyma bands with a central zone of thin-walled cells filled with dark contents that are often missing in abraded cones. Cone-scale vasculature is located in the thin-walled cells of this central zone. There is one large ovule per cone-scale complex with a thick sclerotesta showing a zigzag sclereid pattern. The nucellus is free from the integument except at the chalaza and shows a prominent wavy apex. Megagametophytes and embryos with two cotyledons are poorly preserved but are present in a few seeds. The presence of pollen tubes in the nucellar tissues of...
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