THE SUBTRIBES AND GENERA OF THE TRIBE BROSCINI (COLEOPTERA: CARABIDAE): CLADISTIC ANALYSIS, TAXONOMIC TREATMENT, AND BIOGEOGRAPHICAL CONSIDERATIONS

Abstract

Abstract Phylogenetic relationships of the genera of the tribe Broscini (Carabidae) are postulated based on cladistic methods. Seventy-three morphological characters were examined in 88 species, arranged in 41 genera and subgenera, and with the related tribes Melaenini and Apotomini as outgroups. In all the obtained cladograms, the genera are grouped in five clades recognized as the basis for a new subtribal classification: the Oriental-Palearctic-Neotropical Axonyina, new subtribe (type genus, Axonya Andrewes) with 3 genera; the Holarctic-Oriental subtribe Broscina (type genus Broscus Panzer) with 9 genera; the south temperate Australian-Neotropical Nothobroscina new subtribe (type genus Nothobroscus Roig-Junent and Ball) with 10 genera; the south temperate Neotropical Barypina (type genus Barypus Dejean) with 2 genera; and the south temperate Australian-Neotropical Creobiina (type genus Creobius Guerin-Meneville) with 11 genera. Species groups within diverse genera Barypus, Eurylychnus Bates, and Promec...