Abstract
The difficulty of specifying additive and dominance variation in the presence of epistasis is described. This difficulty can be overcome when the epistasis is at a relatively low level of intensity but when epistasis is the major source of genetic variation no measure of additive or dominance variation is possible. Heterosis in a cross between two inbred lines ofNicotiana rustica was found to be due to dominance opposed to a small extent by epistasis. The misleading nature of another method of partitioning heterosis is pointed out.
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