Abstract
Beyond the better-studied carbohydrates and the macronutrients nitrogen and phosphorus, a remaining 20 or so elements are essential for life and have distinct geographical distributions, making them of keen interest to ecologists. Here, I provide a framework for understanding how shortfalls in micronutrients like iodine, copper, and zinc can regulate individual fitness, abundance, and ecosystem function. With a special focus on sodium, I show how simple experiments manipulating biogeochemistry can reveal why many of the variables that ecologists study vary so dramatically from place to place. I conclude with a discussion of how the Anthropocene's changing temperature, precipitation, and atmospheric CO2levels are contributing to nutrient dilution (decreases in the nutrient quality at the base of food webs).
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