Abstract

Following the disappearance of the seven Buckingham Member (Unit 10) Cypraeidae from the Myakka Lagoon System (Myakka), there was a hiatus represented by two strata at Sarasota (Pinecrest Members 9 & 8) in the early Piacenzian Pliocene, 3.6 million years ago (“mya”), from which no Cypraeidae have been recorded. This was followed by the warmest and most tropical conditions of the entire Pliocene. The sea grass beds and mud flats, which were the preferred habitats of the Pinecrest Cypraeidae, returned in force. The depositional strata which resulted, Pinecrest (Unit 7) and its Kissimmee River Valley Equivalent, were very thick and probably represent a long period of uninterrupted deposition. Eight new species in six genera emerged in Myakka. In the Kissimmee River Valley (Kissimmee), six new species in three genera (one new) emerged as the first and earliest known Cypraeidae species from that area. This represented geographically separate, but parallel, evolutionary tracks. Pinecrest Members 6 & 5 represented a repeat of the conditions experienced during Pinecrest Members 9 & 8, resulting in the disappearance of all Pinecrest (Unit 7) Cypraeidae from Myakka and Kissimmee.

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