Abstract
The chapter on anatomy in the Treatise on Invertebrate Paleontology (Part H, Brachiopoda, revised) (Williams et al., 1997) is the most current and comprehensive treatment that we have of reproduction and development in these animals. My contribution to this short course is a commentary on and addendum to this review. The study of the developmental biology of extant brachiopods describes a large part of their life history and defines several of the parameters that have to be taken into account when thinking about how a given set of genes will make it to the next generation (Havenhand, 1995). Some extant brachiopod genera like Discinisca and Crania (Neocrania) belong to families that first appeared in the fossil record during the Lower Ordovician or, as in the case of Glottidia, to a superfamily that first appeared during the Lower Cambrian. Studies on the development of these extant animals provide a picture of what the development of their Lower Paleozoic ancestors might have been like.
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