The development of terrestrial mollusc faunas in the ‘Rheinland region’ (western Germany and Luxembourg) during the Lateglacial and Holocene

Abstract

A composite faunal history, covering the last ∼13,000 radiocarbon years, has been reconstructed from the few molluscan successions available from the ‘Rheinland’ region. The region, centred on the German Land of Rheinland-Pfalz (but including the Principality of Luxembourg) does not form a discrete biogeographical province but does represent an important link between the better known faunas of central Europe and those of north-west Europe. Open-ground assemblages dominated by Vallonia and Trichia, with few shade-demanding elements, occur throughout the Lateglacial and early Holocene. The Lateglacial is characterised by the presence of Vertigo genesii and Abida secale, depending on whether the depositional environment is wet or dry, respectively. Catholic elements became increasingly important at the start of the Holocene. The establishment of woodland, from approximately 9700 yr BP, is indicated by the expansion of shade-demanding taxa, most notably Carychium tridentatum. Deciduous forest optimum faunas developed by about 7500 yr BP and are characterised by Platyla polita, Perforatella incarnata and Sphyradium doliolum. The modern ranges of these species suggest that during the Holocene Platyla polita, Perforatella incarnata (and possibly Sphyradium doliolum also) colonised northwestern Europe from refugia further east. The reappearance of open-ground faunas from ∼5000 yr BP reflects widespread anthropogenic clearance.