Abstract

This work was supported by a Department of Science and TechnologyCentre of Excellence grant to the FitzPatrick Institute of African Ornithology. R.B.S. was supported by a fellowship from the Leiden Conservation Foundation. This research was approved by SANParks (CONM1182), the Department of Environmental Affairs (RES2013/24, RES2014/83, and RES2015/65) and the animal ethics committee of the University of Cape Town (2013/V3/TC).

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