Abstract

The sustainability of coniferous monoculture plantations is facing challenges with respect to yields, ecology, and biodiversity. Conversion of monocultural coniferous plantations into mixed stands using thinning or direct mixed planting is widely considered to be a key strategy for overcoming these challenges and transforming the characteristics of plantations on a regional scale. Substantial amounts of deadwood may be produced in mixed forests (MFs); this material is important for evaluating and modifying forest management methods, understanding the dynamics of forest stands, and achieving biodiversity conservation. We assessed the quantitative characters and diameter distributions of deadwood in mixed and thinned Chinese fir [Cunninghamia lanceolata (Lamb.) Hook.] forests over one rotation. We used the g(r) function and spatial parameters to analyze the spatial structure of deadwood, and used logistic regression and Hegyi’s competition index (HCI) to explore competition and mortality. Our results indicate that: (1) Chinese fir dominated in all groups of deadwood (snags, broken wood, and fallen wood), and the abundance, volume, and mortality rates of deadwood were much lower in the thinning forest compared to the MF. (2) Later coming populations (LCPs) comprised the majority of the small diameter classes in the thinning forest, but only accounted for a small proportion of the MF. (3) Broken wood in the thinning forest was randomly distributed, while the other types of deadwood were clustered at most spatial scales. In contrast, the spatial patterns in the MF were random at most spatial scales. (4) Total deadwood in both stands was in a status of intermediate and was randomly surrounded by its four nearest neighbors. All types of deadwood were highly mixed in the thinning forest and moderately mixed in the MF. Our case study suggests that thinning and mixing result in different stand development processes and thus influence the type, amount, and structure of deadwood. Thinning significantly reduces competition, which is the main driver of tree mortality. Converting pure Chinese fir plantations into mixed stands by thinning should be taken in future. Understanding tree mortality after conversion is essential to select appropriate silvicultural treatments and achieve ultimately sustainable forest management.

Highlights

  • The conversion of natural forests to plantations has been widely studied, but the findings of these studies cannot be applied to the conversion of plantations to uneven-aged mixed forests (MFs), resulting in an absence of suitable guidelines

  • For sustainable forest management, the issue of timber supply must be resolved; forest biodiversity, ecological functions and benefits, and social services must be attended to, which requires a change in the silvicultural methods used in plantations

  • Some underlying mechanisms of plantations and natural forests maybe beneficial to tree growth (Jelínková et al, 2009; Tarroux and DesRochers, 2011), a small part of retained planted trees was eliminated via long-term competition, highlighting the necessity of thinning

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Summary

Introduction

The dynamics of tree mortality are an important characteristic of forest stands (Harvey and Brais, 2007; Gray and He, 2009; Laarmann et al, 2009; Moussaoui et al, 2020), with implications for forest ecosystem functions (i.e., material cycling and energy flows), forest resilience and stability (Siitonen et al, 2000; Bölöni et al, 2017; Oettel et al, 2020; Zhang et al, 2020). Different types of deadwood are formed following tree death, including standing dead trees (snags), fallen wood (logs), and broken wood (Girona et al, 2019). Natural forests, especially those in temperate regions, have an abundance of deadwood that supports the growth and vitality of retained and regenerating trees (Martin et al, 2020). Data on deadwood in plantations are sparse, and formation processes are poorly understood

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