Abstract

Emergence is examined in populations of Hesperostipa comata, and this grass is compared to individuals, populations, and three additional species of Achnatherum. The Achnatherum species are closely related and more distantly related to H. comata. Emergent properties are assessed through measurements of grass spikelets and the divergence from vectors of isometry as expressed in individuals, populations, species, and species groups. Our results demonstrate that the strongest degrees of emergence are expressed at the levels of individuals, species, and groups of species, with the strongest of all being seen when H. comata is compared with the trio of Achnatherum species; populations express the least emergence. This reflects evolutionary differentiation as expressed through the ontogenetic events that led to the formation of spikelets. It can be related to concepts of local and global time; the flow of energy, matter, and information; and the driving force of the expression of novelty (entropy) in organized and historically constrained biological systems.

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