Abstract

If an animal forages for prey that are distributed in patches, it must use some rule to decide when to leave one patch and go on to another. Charnov's marginal value theorem tells when an animal should leave a patch, but it does not tell how an animal should decide when to leave a patch. This is important if patches vary in quality and the forager must use its experience in a patch to decide when best to leave a patch. In this paper a mathematically tractable stochastic model is presented in which animals forage systematically for prey distributed in patches, and three possible strategies (stopping rules) are considered. The best rule is found and compared with the giving-up time (GUT) rule and a fixed-time rule in which the forager remains until it has exhausted each patch. The GUT rule is better than the fixed-time rule for variable patches, and it is relatively insensitive to environmental changes, but it is not as good as the best rule, which I call the assessment rule. The assessment rule is not only...

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