Abstract

Kin selection, which can lead organisms to behave altruistically to their genetic relatives, works differently when—as is often the case in human societies—altruism can be boosted by social pressure. Here I present a model of social norms enforced by indirect reciprocity. In the model there are many alternative stable allocations of rewards (“distributional norms”); a stable norm is stable in the sense that each player is best off following the norm if other players do the same. Stable norms vary widely in how equally they reward players with unequal abilities. In a population of mixed groups (some group members follow one norm, some follow another, and some compromise) with modest within-group coefficients of relatedness, selection within groups favors those who compromise, and selection between groups favors generous generalized reciprocity rather than balanced reciprocity. Thus evolved social norms can amplify kin altruism, giving rise to a uniquely human mode of kin-based sociality distinct from spontaneous altruism among close kin, or cooperation among non-kin.

Highlights

  • Beating Hamilton’s RuleThis article presents a souped-up version of the theory of kin selection

  • Players differ in their ability to help others (= status); they help and are helped based on their advertised abilities (= assumed roles); and each has an incentive to honestly advertise her ability by following a distributional norm (= shared script)

  • A big motive for developing the theory of kin selection was the problem of altruism: Why does one organism sometimes help another when that help will not be reciprocated? The analogous motive for developing a theory of socially enforced nepotism is the problem of distribution: How are the gains from cooperation distributed? [29] Why does social exchange sometimes shade into uni-directional generosity—balanced reciprocity into generalized reciprocity?

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Summary

Introduction

Beating Hamilton’s RuleThis article presents a souped-up version of the theory of kin selection. Players rely on indirect reciprocity and honest advertising of ability to enforce norms of sharing. The model below, combining game theory and population genetics, in which the starting point is not an individual deciding on her own how much to help her kin, but a group of unequal individuals settling on and enforcing a norm determining who gets how much.

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Conclusion

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