Abstract
Cheloniansliveinasafe-boxthatoffersanef-Þcientprotectionagainstmostpredators,atleastin adults and sub-adults (Wilbur and Morin,1988). Nevertheless, in hatchlings, and to alesser extent, in juveniles the shell is soft and/ornot solid enough to resist to many predatorssuch as monitors, dogs, or birds for example(DouglasandWinegarner,1977;FitzpatrickandWoolfenden, 1978; Branch and Els, 1990; Gef-fen and Mendelssohn, 1997; Keller et al., 1998;Hill, 1999; Lagarde et al., 2001; Boarman,2002). In an open landscape, the predation onthese fragile tortoises can be severe; sometimesleaving strong marks on the population dynam-ics (Boarman, 2002). In April 2003, during aÞeld study on the ecology of Testudo graecagraeca in Morocco (Essaouira region, 31 23 N,9 42 W), we found several hatchling tortoisesimpaled on spiny shrubs ( Argania spinosa )ina typical shrike manner (Cramps and Perrins,1993). Therefore, several observations of shrike(Lanius excubitor ) were made in the study areaand even on the involved shrubs. We searchedintensively the study site and the surroundingarea (the total prospected area was 50 ha) anddiscovered a total of 25 impaled tortoises in 3spiny shrubs (respectively 17, 7 and 1 tortoisespershrub).WithanaverageShriketerritorysizeof 75 ha (Cramps and Perrins, 1993), such pre-dation may be due only to one Shrike breed-
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