Abstract
Summary Anatomical aspects of ovule, embryo and endosperm development are described for the zygote to cotyledon embryo stages in A. demavendicus and A. latifolius . Nucellar cells disappear during an early stage of development. The inner integument differentiates into an endothelium and an exterior layer of cells at the zygote stage. The primary endosperm nucleus divides before the zygote nucleus, forming a nuclear endosperm. In A. latifolius the polar nuclei fuse immediately after cellularization of the embryo sac, whereas in A. demavendicus they remain separated until fertilization. Endosperm cellularization begins when the embryo has developed the early globular stage in A. latifolius and the late globular stage in A. demavendicus . The inner integument is degraded at early heart stage in A. latifolius and at late heart stage in A. demavendicus , then the outer integument adjoins the embryo sac. The outer integument consists of three different tissues at this stage. As expected, the studied species which both belong to the same section of Astragalus , show major similarities in seed development. The remarked differences should be considered as variations among the species of this section.
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