Abstract
New morphological information about the female and larva of Psychoglypha mono Denning supports congeneric relationship of this species with Desmona bethula Denning; P. mono is accordingly transferred to Desmona as a second species. Larvae of both species of Desmona live in small spring-fed channels of moist alpine meadow habitats in western North America; those of D. mono also have been found in the littoral zone of small alpine lakes. Larvae burrow into sand-gravel substrates for pupation in late summer, emerging as adults in September and October. Diagnostic characters are provided for adults and larvae of the two species of Desmona ; a summary of their geographic distribution is included. Desmona and the Holarctic Chilostigma are sister groups, forming with Brachypsyche (Palearctic) and Psychoglypha (Nearctic) a closely related complex within the limnephiline tribe Chilostigmini. Because Psychoglypha mono was the type species of the monobasic subgenus P. (Monophylax) Nimmo, a consequence of its transfer to Desmona is that Monophylax becomes a junior synonym of Desmona Denning.
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