Abstract
Population models have suggested that host regulation by natural enemies is achieved by the densitydependent response of natural enemies to temporal and spatial changes in host density. However, surveys of insect host-parasitoid systems (Morrison and Strong 1980, Dempster 1983, Stiling 1987) indicate that few field studies provide unambiguous evidence of densitydependent relationships. It has been argued that density dependence may be obscured by various behavioral and stochastic elements involved in the interaction, rendering our methods for its detection inadequate (Morrison and Strong 1980, Stiling and Strong 1982, Hassell 1985, Lessels 1985, Morrison 1986, Stiling 1987, Price 1988). Hassell (1985) has suggested that the regulatory role of natural enemies may be detected by examining the relationship between the per capita searching efficiency and parasitoid density. The per capita searching efficiency is calculated as:
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