Abstract

Isaza,Chaenogobius isaza, is a small gobiid fish endemic to Lake Biwa. It lives offshore throughout almost the entire year, showing a remarkable diel vertical migration. In early spring, males and females migrate to lake shore to spawn under stones. Spawning season is limited to a very short span of time in early spring, late April to early May. During this short spawning season, the male is supposed to have only 2 brood cycles at maximum. At each brood cycle, the male is strictly monogynous, never accepting additional females. Males therefore show a marked mate choice, choosing a larger female regardless of the size of the male himself. Females also choose larger males. However, males are supposed to not waste time in male-male fighting in the natural spawning area. That this very short spawning season and its occurrence in early spring is primarily to avoid interspecific competition with another littorally reproducing goby Yoshinobori,Rhinogobius brunneus, is experimentally demonstrated.

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