Abstract

<p>The modelled global vegetation for the end of the 21st century is currently is insufficiently constrained<br>by climate models. A significant proportion of that uncertainty has been attributed to the limitations<br>of current Dynamic Global Vegetation Models (DGVMs), and the misrepresentation of mortality, dis-<br>turbance and regrowth within forests. Improving the simulation of the underlying processes of de-<br>mographic change is of primary importance in the development of predictors of future climate.</p><p>Here we present the Robust Ecosystem Demography (RED), a new dynamical vegetation model which<br>simulates the size-structure of forests by partitioning the population of a Plant Functional Type (PFT)<br>into mass classes. Allometric scaling of mortality and growth across mass classes allows for a variety<br>of complex demographic processes to be captured, such as disturbances and regrowth. Competition<br>among PFTs is done purely through restricting the recruitment of new vegetation to unshaded space.<br>RED represents a reduction of complexity from more numerically unwieldy cohort DGVMs which<br>simulate both size and patch dimensions. The limited number of dimensions and simple competitive<br>regime allows the equilibrium state to be solved for analytically, providing two potential functions - (i)<br>Avoiding-spinning up by providing an equilibrium state for intilisation. (ii) Insights into the demog-<br>raphy of vegetated areas, arising from parameter tuning to fit observation, such as coverage or carbon<br>mass. When paired with a rate of mortality or carbon assimilate rate gives, respectfully, required as-<br>similate or mortality rates.<br>We demonstrate the model functionality using offline UKESM PFT carbon assimilate rates, paired<br>with observed vegetation cover from the ESA LC_CCI datasets for the 9 different PFTs. From this<br>dataset we calibrate a novel global equilibrium mortality map for each PFT and show the competitive<br>and successional behaviour of dynamical runs with convergence to the fitted equilibrium. Finally, we<br>explore underlying ecological questions that emerge from the equilibrium solutions.</p>

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