Abstract

Piñon (Pinus spp. L.)–juniper (Juniperus spp. L.) woodlands’ historical stand structures were recreated to provide reference conditions and document long-term changes in the Sky Islands of the Davis Mountains, Texas. Restoration of these isolated woodlands requires insights into the range of variability in current and historical stand structures, as well as an understanding of the spatiotemporal establishment and recruitment patterns of tree species. With drastic changes in forests and woodlands of the Southwestern United States widely reported, the main objective of this study was to reconstruct woodland tree temporal and spatial establishment patterns. A stratified random sampling approach was used to select two study sites each of 3600 m2 in area. Within each site, all individual trees were mapped, measured, and cored for age determination. Age and tree location data were used to recreate the spatiotemporal patterns of tree species establishment and recruitment. Increments in density of both Mexican piñon (Pinus cembroides var. cembroides Zucc.) and alligator juniper (Juniperus deppeana var. deppeana Steud.) reached 422 trees ha−1 in the 115-year period between 1890 and 2005; a yearly increment of 4 trees ha−1 that did not reflect a rapid rate of change in these piñon–juniper woodlands. Age distributions reflected the multi-cohort nature of these woodlands, and spatial autocorrelation measures were useful in the objective delineation of these cohorts. Temporal and functional niche differentiation of juniper was reflected in the development pattern where alligator juniper served as a pioneer species, exhibited a longer period of substantial recruitment, and had greater recruitment rates than that of Mexican piñon. Recruitment of Mexican piñon and alligator juniper occurred in an episodic fashion, with the majority of recruits being acquired between 1890 and 1949.

Highlights

  • Reconstructive studies enable the assessment of long-term changes and ecosystem dynamics, aid in the interpretation of ecosystem disturbance history, and are essential in guiding conservation and management goals [1,2]

  • Alligator juniper density increased from 33 trees/ha in the 1890s to 274 trees/ha in 2005, whereas Mexican piñon density increased from 7 trees/ha in the 1890s to 188 trees/ha in 2005 (Figure 1)

  • Stand structure reconstruction in this piñon–juniper woodland revealed substantial increments in density, similar to those reported in the Great Basin and Northern Arizona [15,20]

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Summary

Introduction

Reconstructive studies enable the assessment of long-term changes and ecosystem dynamics, aid in the interpretation of ecosystem disturbance history, and are essential in guiding conservation and management goals [1,2]. Examination of changes in forest and woodland stand structure over time can be accomplished through the use of permanent plots that are monitored for tree recruitment, growth, and mortality at regular intervals. This approach may be limited in application due to inherent costs, as well as changes in land ownership. An alternative approach is to determine age structure retrospectively using dead and live trees [3]. The use of static age structures (at one point in time) in the interpretation of stand dynamics has been scrutinized, .

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