Abstract
Fifty plots on grassland on well-drained soil were investigated for land use history and analysed for the species composition of vascular plants, macrofungi, bryophytes, lichens, hepatics and for 20 environmental variables. A DCA ordination showed an overall influence of land use history on species composition. DCA axis 1 was strongly correlated with species diversity of vascular plants. High species diversity is proposed to result from the combined action of high pH, high CaCO 3 content, high age of turf, grazing and absence of fertilization. A DCCA ordination lead to the proposal of 22 species as indicators of old, unameliorated grasslands in Denmark.
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