Abstract
In the course of evolution, the adaptation of aquatic organisms to terrestrial life has caused many changes in their morphology, physiology and behaviour. As a result of the invasion of land, large groups of species evolved entirely independently of the aquatic environment. Among the aquatic invertebrates a small group of decapods crustaceans, living where the sea and land merges, are unique in that they are really amphibious, living part of their lives on the land and the other part in the water. Some grapsid crabs belong here in addition to several species of the ghost crabs (Ocypode), species of Uca, Scopimera and Dotilla, hermit crabs (Coenobita and Virgo) and others. Within the Red Sea many of them are found along sandy beaches and mangrove zones, and thus characterize widespread communities (Fishelson, 1971).
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