Abstract
Although studies of geographic variation are relatively common, the vast majority have dealt with morphometric or meristic traits (13, 120, 297, 327), allozymes (150, 233, 327), or most recently, mitochondrial DNA (9). The relative rarity of studies of geographic variation in traits (see 42, 245, 312 for reviews of early studies) is somewhat surprising, considering that a major focus within vertebrate physiological ecology, comparative physiology, environmental physiology, and biophysical ecology is the study of adaptation (14, 15, 44, 75, 102, 112, 156, 181, 244, 246, 298, 299). Of course, populations may show differentiation on other than large geographic scales, such as microgeographic (150, 188, 213, 233, 290),
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